Mimid
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__NOTOC__ The mimids are the
New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...
family of
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
birds, Mimidae, that includes thrashers,
mockingbird Mockingbirds are a group of New World passerine birds from the family (biology), family Mimidae. They are best known for the habit of some species Mimicry, mimicking the songs of other birds and the sounds of insects and amphibians, often loudly ...
s, tremblers, and the New World catbirds. As their name (
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
for "mimic") suggests, these
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s are notable for their vocalization, especially some species' remarkable ability to mimic a wide variety of birds and other sounds heard outdoors. They are commonly referred to as mimic thrushes but are not, in fact, thrushes (which are members of family Turdidae).


Description

There are over 30
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of mimids in two larger and some ten small or
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
. They tend toward dull grays and browns in their appearance, though a few are black or blue-gray, and many have red, yellow, or white irises. They range from 20 to 33 centimetres in length, and 36 to 56 grams in weight. Many mimids have a rather thrush-like pattern: brown above, pale with dark streaks or spots below. They tend to have longer tails than thrushes (or the bigger
wren Wrens are a family, Troglodytidae, of small brown passerine birds. The family includes 96 species and is divided into 19 genera. All species are restricted to the New World except for the Eurasian wren that is widely distributed in the Old Worl ...
s, which they also resemble) and longer bills that in many species curve downward.Clement; Peter; Perrins, Christopher (2003). Mockingbirds. ''In:'' Perrins, Christopher (ed.): ''The Firefly Encyclopedia of Birds'': 534–535. Firefly Books. They have long, strong legs (for passerines) with which many species hop through undergrowth searching for
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s and fruits to eat. Their habitat varies from forest undergrowth to scrub, high-altitude grasslands, and deserts. The two tremblers live in the atypical habitat of
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s in the
Lesser Antilles The Lesser Antilles is a group of islands in the Caribbean Sea, forming part of the West Indies in Caribbean, Caribbean region of the Americas. They are distinguished from the larger islands of the Greater Antilles to the west. They form an arc w ...
, and the brown trembler has the particularly atypical behavior of foraging while clinging to tree trunks. All known species build somewhat messy, bulky twig nests in dense growth, which are in most species on the ground or no more than 2 meters up. They usually lay 2 to 5 eggs that hatch in 12 or 13 days, which is also the length of time the chicks stay in the nest. Breeding usually starts in the spring or early in the rainy season, and many species can have two or even three broods per year. Most failures to fledge young are due to predation. Pairs often stay together for more than one breeding season.


In the history of science

Contrary to often-held belief, the '' Nesomimus'' mockingbirds may have played at least as great a role as
Darwin's finches Darwin's finches (also known as the Galápagos finches) are a group of about 18 species of passerine birds. They are well known for being a classic example of adaptive radiation and for their remarkable diversity in beak form and function. They ...
in inspiring Darwin's work on his theory of evolution.


Systematics


Outside the family

Phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analyses have shown that mimids are most closely related to
starling Starlings are small to medium-sized passerine (perching) birds known for the often dark, glossy iridescent sheen of their plumage; their complex vocalizations including mimicking; and their distinctive, often elaborate swarming behavior, know ...
s. Sibley, Charles Gald & Monroe, Burt L. Jr. (1990). ''Distribution and taxonomy of the birds of the world: A Study in Molecular Evolution''. Yale University Press, New Haven, CT. Zuccon, Dario; Cibois, Anne; Pasquet, Eric & Ericson, Per G.P. (2006). Nuclear and mitochondrial sequence data reveal the major lineages of starlings, mynas and related taxa. ''Molecular Phylogenetics and Evolution'' 41(2): 333–344. (HTML abstract) These and oxpeckers (and the Philippine creepers if they are not outright but highly
apomorph In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ha ...
ic starlings) form a group of Muscicapoidea which originated probably in the
Early Miocene The Early Miocene (also known as Lower Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages: the Aquitanian age, Aquitanian and Burdigalian stages. The sub-epoch lasted from 23.03 ± 0.05 annum, Ma to ...
– very roughly 25–20 mya – somewhere in
East Asia East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...
. This is evidenced by the Asian-SW Pacific distribution of the most basal starlings (and Philippine creepers) and the North American range of the basal mimids. They are sometimes united with the starlings in the Sturnidae as a
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Mimini as proposed by Sibley & Monroe (1990). This makes the expanded Sturnidae a rather noninformative group and is probably due to the methodological drawbacks of their DNA-DNA hybridization technique.


Within the family

The mockingbirds with some thrashers seem to form one major
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
, while the two other groups and the remaining thrashers seem to form another, but the basal branching pattern is not well resolved. The tremblers, again, are a
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
lineage. The latter, however, are embedded in a
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
catbird-
Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...
thrasher assemblage which consists of many rather basal lineages. For detailed information on the
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
ary relationships of the different mimid lineages, see their articles.
Mockingbird Mockingbirds are a group of New World passerine birds from the family (biology), family Mimidae. They are best known for the habit of some species Mimicry, mimicking the songs of other birds and the sounds of insects and amphibians, often loudly ...
s: * Genus '' Mimus'' – typical mockingbirds (some 10 species, includes ''Mimodes'') * The former genus '' Nesomimus'', now part of ''Mimus''American Ornithologists' Union, "Changes since 1 March 2005"
/ref> – mockingbirds of the
Galápagos Islands The Galápagos Islands () are an archipelago of volcanic islands in the Eastern Pacific, located around the equator, west of the mainland of South America. They form the Galápagos Province of the Republic of Ecuador, with a population of sli ...
(4 species) * Genus '' Melanotis'' – blue mockingbirds (2 species) New World catbirds: * Genus '' Dumetella'' – grey catbird * Genus '' Melanoptila'' – black catbird Thrashers: * Genus '' Oreoscoptes'' – sage thrasher * Genus '' Toxostoma'' – typical thrashers (10 species) * Genus '' Ramphocinclus'' – white-breasted thrasher * Genus '' Allenia'' – scaly-breasted thrasher (formerly in ''Margarops'') * Genus '' Margarops'' – pearly-eyed thrasher Tremblers * Genus '' Cinclocerthia'' (2 species)


References


External links


Mimid videos
on the Internet Bird Collection {{Authority control Birds of the Americas Taxa named by Charles Lucien Bonaparte