Millettia Sutherlandii
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''Millettia'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
flowering plants Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
. It consists of about 169
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of shrubs,
liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...
s or trees, which are native to
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
regions of sub-Saharan Africa, the Indian subcontinent, Indochina, southern China,
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms. It is a phytogeographical floristic region in the Paleotropical kingdom. It was first recognized as a distinct region ...
, and
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
. Typical habitats include tropical rain forest and seasonally-dry lowland and upland forest and forest margins, woodland, thicket, wooded grassland, and secondary vegetation.


Description

In 1834, in ''Prodromus Florae Peninsulae Indiae Orientalis''
Robert Wight Robert Wight (6 July 1796 – 26 May 1872) was a Scottish surgeon in the East India Company, whose professional career was spent entirely in southern India, where his greatest achievements were in botany – as an economic botanist and leading ...
and
George Arnott Walker-Arnott George Arnott Walker Arnott of Arlary (6 February 1799 – 17 April 1868) was a Scottish botanist. He collaborated with botanists from around the world and served as a Regius Professor of Botany (Glasgow), regius professor of botany at the Uni ...
describe ''Millettia'' as:
Calyx CALYX, Inc. is a non-profit publisher of art and literature by women founded in 1976 based in Corvallis, Oregon. CALYX publishes both '' CALYX, A Journal of Art and Literature by Women'' twice a year and CALYX Books, which publishes one to three ...
cup-shaped, lobed or slightly toothed. Corolla
papilionaceous Papilionaceous flowers (from Latin: ''papilion'', a butterfly) are flowers with the characteristic irregular and butterfly-like corolla found in many, though not all, plants of the species-rich Faboideae subfamily of legumes. Tournefort sugge ...
: vexillum recurved, broad, emarginate, glabrous or silky on the back. Stamens diadelphous (9 and 1), the tenth quite distinct.
Legume Legumes are plants in the pea family Fabaceae (or Leguminosae), or the fruit or seeds of such plants. When used as a dry grain for human consumption, the seeds are also called pulses. Legumes are grown agriculturally, primarily for human consum ...
flat, elliptic or lanceolate, pointed, coriaceous, thick margined, wingless indehiscent, 1-2 seeded: valves closely cohering with each other all round the seeds and between them. Twining or arboreous. Leaves very large, unequally pinnated: leaflets opposite, with a setaceous partial stipule at the base of each partial
petiole Petiole may refer to: *Petiole (botany), the stalk of a leaf, attaching the blade to the stem *Petiole (insect anatomy) In entomology, petiole is the technical term for the narrow waist of some hymenopteran insects, especially ants, bees, and ...
.
Raceme A raceme () or racemoid is an unbranched, indeterminate growth, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are ...
s axillary, more or less branched and compound. Flowers pretty large, purplish, pedicelled on shortish diverging partial peduncles.


Etymology

Long known to residents of the
Indies The East Indies (or simply the Indies) is a term used in historical narratives of the Age of Discovery. The ''Indies'' broadly referred to various lands in the East or the Eastern Hemisphere, particularly the islands and mainlands found i ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, and
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, this species has had many traditional names. One of the oldest references in
traditional Chinese medicine Traditional Chinese medicine (TCM) is an alternative medicine, alternative medical practice drawn from traditional medicine in China. A large share of its claims are pseudoscientific, with the majority of treatments having no robust evidence ...
is in ''
Bencao Gangmu Shiyi Bencao may refer to: * Xinxiu bencao The ''Xinxiu bencao'' (), also known as the ''Tang bencao'' (), is a Chinese pharmacopoeia written in the Tang dynasty by a team of officials and physicians headed by editor-in-chief . It borrowed heavily fr ...
'' ("Supplement to
Compendium of Materia Medica The ''Bencao gangmu'', known in English as the ''Compendium of Materia Medica'' or ''Great Pharmacopoeia'', is an encyclopedic gathering of medicine, natural history, and Chinese herbology compiled and edited by Li Shizhen and published in th ...
") where is called jixueteng. The Chinese name literally translates to "stem of chicken's blood" which refers to the red resin present in the stems of several climbing legume shrubs. In the 1820s-1830s Charles Millett, a plant collector and an official with the
East India Company The East India Company (EIC) was an English, and later British, joint-stock company that was founded in 1600 and dissolved in 1874. It was formed to Indian Ocean trade, trade in the Indian Ocean region, initially with the East Indies (South A ...
, collected many samples of Millettia while living in
Canton Canton may refer to: Administrative divisions * Canton (administrative division), territorial/administrative division in some countries * Township (Canada), known as ''canton'' in Canadian French Arts and entertainment * Canton (band), an It ...
and
Macao Macau or Macao is a special administrative region of the People's Republic of China (PRC). With a population of about people and a land area of , it is the most densely populated region in the world. Formerly a Portuguese colony, the ter ...
. He sent them to the
University of Glasgow The University of Glasgow (abbreviated as ''Glas.'' in Post-nominal letters, post-nominals; ) is a Public university, public research university in Glasgow, Scotland. Founded by papal bull in , it is the List of oldest universities in continuous ...
's Botanical Garden. In 1834,
Robert Wight Robert Wight (6 July 1796 – 26 May 1872) was a Scottish surgeon in the East India Company, whose professional career was spent entirely in southern India, where his greatest achievements were in botany – as an economic botanist and leading ...
and
George Arnott Walker-Arnott George Arnott Walker Arnott of Arlary (6 February 1799 – 17 April 1868) was a Scottish botanist. He collaborated with botanists from around the world and served as a Regius Professor of Botany (Glasgow), regius professor of botany at the Uni ...
, both Scottish botanists, published ''Prodromus Florae Peninsulae Indiae Orientalis'' where the genus ''Millettia'' is first mentioned. The authors named the genus after Charles Millett, incorrectly referring to him as Dr. Charles Millett. Charles Millett of the East India Company has often been confused with Charles Millet, a French
ichthyologist Ichthyology is the branch of zoology devoted to the study of fish, including bony fish (Osteichthyes), cartilaginous fish (Chondrichthyes), and jawless fish (Agnatha). According to FishBase, 35,800 species of fish had been described as of March 2 ...
, who was active around the same time. In addition J. A. Millet, a French botanist from the 18th century, is often misattributed as the source. Robert Sweet states that the genus ''
Pongamia ''Pongamia pinnata'' is a species of tree in the pea family, Fabaceae, native to eastern and tropical Asia, Australia, and the Pacific islands. It is the sole species in genus ''Pongamia''. It is often known by the synonym ''Millettia pinnata''. ...
'' comes from the
Malabar region The Malabar Coast () is the southwestern region of the Indian subcontinent. It generally refers to the western coastline of India stretching from Konkan to Kanyakumari. Geographically, it comprises one of the wettest regions of the subcontin ...
in India and is derived from the local word Pongam (most likely from the
Malayalam Malayalam (; , ) is a Dravidian languages, Dravidian language spoken in the Indian state of Kerala and the union territories of Lakshadweep and Puducherry (union territory), Puducherry (Mahé district) by the Malayali people. It is one of ...
language). ''Pongamia'' had often been misattributed to Vent. (1803), but it was preceded by "Pongam Adans. (1763)", "Galedupa Lam. (1788)", and "Pungamia Lam. (1796)" and in accordance with the 1994 Tokyo Code of the
International Code of Botanical Nomenclature The ''International Code of Nomenclature for algae, fungi, and plants'' (ICN or ICNafp) is the set of rules and recommendations dealing with the formal botanical names that are given to plants, fungi and a few other groups of organisms, all tho ...
, the correct citation was established as "Pongamia Adans. (1763)".Plant Name Details - Leguminosae Pongamia Vent.
/ref> In 1981 a proposal to conserve the genus ''Millettia'' and reject the genus ''Pongamia'' was proposed in the journal
Taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
and was ratified in 1988. Most of the species formerly classed in ''Pongamia'' are now included in ''Millettia'',Smithsonian National Museum of Natural History - Proposal No. 549
/ref> with the exception of ''
Pongamia pinnata ''Pongamia pinnata'' is a species of tree in the pea family, Fabaceae, native to eastern and tropical Asia, Australia, and the Pacific islands. It is the sole species in genus ''Pongamia''. It is often known by the synonym ''Millettia pinnata''. ...
''.Wendy E. Cooper, Darren M. Crayn, Frank A. Zich, Rebecca E. Miller, Melissa Harrison, Lars Nauheimer "A review of Austrocallerya and Pongamia (Leguminosae subfamily Papilionoideae) in Australia, and the description of a new monotypic genus, Ibatiria," ''Australian Systematic Botany'', 32(4), 363-384, (29 August 2019) https://doi.org/10.1071/SB18039


Uses

Species are used locally as fuelwood, fish poisons, insecticides, medicine, ornamentals, and nitrogen fixers for soil rehabilitation in
agroforestry Agroforestry (also known as agro-sylviculture or forest farming) is a land use management system that integrates trees with crops or pasture. It combines agricultural and forestry technologies. As a polyculture system, an agroforestry system c ...
. Several species are used for timber, including the African species wenge ('' M. laurentii'' ) and panga panga or mpande ('' M. stuhlmannii'' ). The timber is used for flooring, furniture, cabinet work, construction, veneers, joinery, and agricultural implements.


Species

Selected species include: * '' Millettia aurea'' * '' Millettia australis'' - Samson's sinew (Norfolk Island) * ''
Millettia brandisiana ''Millettia'' is a genus of flowering plants in the family Fabaceae. It consists of about 169 species of shrubs, lianas or trees, which are native to tropical and subtropical regions of sub-Saharan Africa, the Indian subcontinent, Indochina, sout ...
'' * ''
Millettia bussei ''Millettia bussei'' is a species of plant in the family Fabaceae. It is found in Mozambique and Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered ...
'' * ''
Millettia capuronii ''Millettia capuronii'' is a species of plant in the family Fabaceae. It is found only in Madagascar. The Latin specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclat ...
'' * ''
Millettia conraui ''Millettia conraui'' is a species of plant in the family Fabaceae. It is found in Cameroon and Nigeria. It is threatened by habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is n ...
'' * '' Millettia diptera'' Gagnep. * ''
Millettia duchesnei ''Millettia duchesnei'' is a large forest liana in the genus ''Millettia''. The rotenoids elliptol, , , , , , and the flavanone eriodictyol Eriodictyol is a bitter-masking flavanone, a flavonoid extracted from yerba santa ('' Eriodictyon cal ...
'' * ''
Millettia elongatistyla ''Millettia elongatistyla'' is a species of legume in the family Fabaceae. It is found only in Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered b ...
'' * '' Millettia eriocarpa'' * '' Millettia galliflagrans'' * ''
Millettia grandis ''Millettia'' is a genus of flowering plants in the family Fabaceae. It consists of about 169 species of shrubs, lianas or trees, which are native to tropical and subtropical regions of sub-Saharan Africa, the Indian subcontinent, Indochina, sout ...
'' – Umzimbeet * '' Millettia hitsika'' * '' Millettia lacus-alberti'' * ''
Millettia laurentii ''Millettia'' is a genus of flowering plants in the family Fabaceae. It consists of about 169 species of shrubs, lianas or trees, which are native to tropical and subtropical regions of sub-Saharan Africa, the Indian subcontinent, Indochina, sout ...
'' – Wengé * '' Millettia macrophylla'' * '' Millettia micans'' * '' Millettia mossambicensis'' * ''
Millettia nathaliae ''Millettia nathaliae'' is a species of plant in the family Fabaceae. It is found only in Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peri ...
'' * '' Millettia nigrescens'' Gagnep. * ''
Millettia nitida ''Callerya nitida'' is a species of flowering plant in the family Fabaceae, native to south-central and southeast mainland China, Hainan and Taiwan. It was first described by George Bentham in 1842 as ''Millettia nitida''. ''Callerya nitida'' p ...
'' * ''
Millettia orientalis ''Millettia orientalis'' is a species of plant in the family Fabaceae. It is found only in Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller per ...
'' * ''
Millettia peguensis ''Millettia peguensis'', the Moulmein rosewood (Bengali: তূমা Tuma), is a legume tree species in the genus ''Millettia''. It is native to Lower Burma Myanmar, officially the Republic of the Union of Myanmar; and also referred t ...
'' * ''
Millettia psilopetala ''Millettia psilopetala'' is a species of legume in the family Fabaceae. It is found in Democratic Republic of the Congo and Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the e ...
'' * '' Millettia pterocarpa'' * '' Millettia pubinervis'' Kurz * '' Millettia puerarioides'' Prain * '' Millettia richardiana'' * '' Millettia rhodantha'' * ''
Millettia sacleuxii ''Millettia'' is a genus of flowering plants in the family Fabaceae. It consists of about 169 species of shrubs, lianas or trees, which are native to tropical and subtropical regions of sub-Saharan Africa, the Indian subcontinent, Indochina, sout ...
'' * '' Millettia schliebenii'' * ''
Millettia semseii ''Millettia semseii'' is a species of legume in the family Fabaceae. It is endemic to Tanzania. It has been assessed as " vulnerable" in the IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Spec ...
'' * '' Millettia sericantha'' * ''
Millettia stuhlmannii ''Millettia stuhlmannii'', commonly known as panga panga, is a well-known species of timber tree that is native to the southeastern Afrotropics. The wood of the tropical species '' M. laurentii'' has similar qualities and uses, but is slightly da ...
'' – Panga panga * '' Millettia sutherlandii'' * '' Millettia taolanaroensis'' * '' Millettia thonningii'' * '' Millettia usaramensis'' – Lesser millettia * '' Millettia utilis'' * '' Millettia warneckei'' * ''
Millettia xylocarpa ''Millettia xylocarpa'' is a species of plant in the family Fabaceae. It is a tree native to Indochina, Peninsular Malaysia, Sumatra, Java, Borneo, and the Philippines. It is threatened by habitat loss Habitat destruction (also termed habit ...
''


References

{{Taxonbar, from=Q2628493 Fabaceae genera Taxa named by Robert Wight Taxonomy articles created by Polbot Paleotropical flora Taxa named by George Arnott Walker Arnott