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Milacidae is a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of air-breathing,
keel The keel is the bottom-most longitudinal structural element of a watercraft, important for stability. On some sailboats, it may have a fluid dynamics, hydrodynamic and counterbalancing purpose as well. The keel laying, laying of the keel is often ...
ed, land
slug Slug, or land slug, is a common name for any apparently shell-less Terrestrial mollusc, terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced ...
s. These are shell-less
terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
s in the superfamily
Parmacelloidea Parmacelloidea is a Taxonomic rank, superfamily of air-breathing land slugs, terrestrial molluscs, terrestrial pulmonate gastropod Mollusc, mollusks in the clade Stylommatophora and the informal group Pulmonata. These are limacoid or keelback slu ...
. This family has no subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).


Distribution

The distribution of the Milacidae includes the western
Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...
region.


Anatomy

In the Milacidae family, the number of
haploid Ploidy () is the number of complete sets of chromosomes in a cell (biology), cell, and hence the number of possible alleles for Autosome, autosomal and Pseudoautosomal region, pseudoautosomal genes. Here ''sets of chromosomes'' refers to the num ...
chromosome A chromosome is a package of DNA containing part or all of the genetic material of an organism. In most chromosomes, the very long thin DNA fibers are coated with nucleosome-forming packaging proteins; in eukaryotic cells, the most import ...
s ranges from 31 to 35 (according to the values in this table).Barker G. M.: Gastropods on Land: ''Phylogeny, Diversity and Adaptive Morphology''. in Barker G. M. (ed.):
The biology of terrestrial molluscs
'. CABI Publishing, Oxon, UK, 2001, . 1-146, cited pages: 139 and 142.


Genera

Genera within the family Milacidae include: * '' Milax'' Gray, 1855 - type genus * ''
Tandonia ''Tandonia'' is a genus of air-breathing, keeled, land slugs. These are shell-less terrestrial gastropod mollusks in the family Milacidae. Distribution The distribution of the genus ''Tandonia'' includes Europe to North Africa and Asia Minor. ...
'' Lessona & Pollonera, 1882


Cladogram

The following
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
shows the
phylogenic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...
relationships of this family to the other families in the
limacoid clade The Limacoidei is a taxonomic infraorder of air-breathing land snails, semislugs and slugs, terrestrial pulmonate gastropod molluscs in the suborder Helicina. Distribution The original ancestral area of limacoid families is thought to be ...
:Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". ''
Journal of Biogeography The ''Journal of Biogeography'' is a monthly peer-reviewed scientific journal that was established in 1974. It covers aspects of spatial, ecological, and historical biogeography. The founding editor-in-chief was David Watts, followed by John Flen ...
'' 27(2): 379-390.
JSTOR


References


Further reading

* Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". ''Ruthenica'', Supplement 2. 1309-1466. * Wiktor A. (1987). "Milacidae (Gastropoda, Pulmonata) — systematic monograph". ''Annales Zoologici'', Warszawa 41: 153-319.


External links

* {{Taxonbar, from=Q1145729