Micrelaps Vaillanti
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''Micrelaps vaillanti'', also known commonly as the black-headed micrelaps or the Somali two-headed snake, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of mildly
venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...
rear-fanged A snake skeleton consists primarily of the skull, vertebrae, and ribs, with only vestigial remnants of the limbs. Skull The skull of a snake is a very complex structure, with numerous joints to allow the snake to swallow prey far larger than it ...
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Micrelapidae :''Common names: two-headed snakes.'' ''Micrelaps'' is a genus of rear-fanged mildly venomous snakes. It is the only genus in the family Micrelapidae. The genus is native to Africa and the Middle East, and there are four species that are recogn ...
. The species is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
.


Etymology

The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''vaillanti'', is in honor of
French French may refer to: * Something of, from, or related to France ** French language, which originated in France ** French people, a nation and ethnic group ** French cuisine, cooking traditions and practices Arts and media * The French (band), ...
herpetologist Herpetology (from Ancient Greek ἑρπετόν ''herpetón'', meaning "reptile" or "creeping animal") is a branch of zoology concerned with the study of amphibians (including frogs, salamanders, and caecilians (Gymnophiona)) and reptiles (in ...
Léon Vaillant Léon Louis Vaillant (; 11 November 1834 – 24 November 1914) was a French zoology, zoologist. He is most famous for his work in the areas of herpetology, malacology, and ichthyology. In 1854 he graduated from the Arras College, College d'Ar ...
.


Geographic range

''M. vaillanti'' is found in
Ethiopia Ethiopia, officially the Federal Democratic Republic of Ethiopia, is a landlocked country located in the Horn of Africa region of East Africa. It shares borders with Eritrea to the north, Djibouti to the northeast, Somalia to the east, Ken ...
,
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
,
Somalia Somalia, officially the Federal Republic of Somalia, is the easternmost country in continental Africa. The country is located in the Horn of Africa and is bordered by Ethiopia to the west, Djibouti to the northwest, Kenya to the southwest, th ...
, eastern
Sudan Sudan, officially the Republic of the Sudan, is a country in Northeast Africa. It borders the Central African Republic to the southwest, Chad to the west, Libya to the northwest, Egypt to the north, the Red Sea to the east, Eritrea and Ethiopi ...
,
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
, and
Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...
.


Habitat

The preferred natural
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s of ''M. vaillanti'' are
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
and
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
, at altitudes from sea level to .


Description

''M. vaillanti'' is brown
dorsally Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provi ...
, with the center of each
dorsal scale In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...
grayish white. The
ventrals In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...
are brown in the middle, and whitish on the sides. Adults may attain a total length of , with a tail long. The dorsal scales are smooth, without pits, and are arranged in 15 rows at midbody (in 17 rows on the neck). The ventrals number 171–203. The anal plate is divided, and the
subcaudals In snakes, the subcaudal scales are the enlarged plates on the underside of the tail. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Comstock Publishing Associates, a Division of Cornell University Press. (7t ...
are also divided. The head is very flattened. The
rostral Rostral may refer to: Anatomy * Rostral (anatomical term), situated toward the oral or nasal region * Rostral bone, in ceratopsian dinosaurs * Rostral organ, of certain fish * Rostral scale The rostral scale, or rostral, in snakes and other sca ...
is large, twice as broad as deep, the portion visible from above about as long as its distance from the frontal. The
internasals In snakes, the internasal scales are those on top of the head between the scales that surround the nostrils. Wright AH, Wright AA (1957). ''Handbook of Snakes of the United States and Canada''. Ithaca and London: Comstock Publishing Associates. (7 ...
are twice as broad as long, and twice as long as the prefrontals. The frontal is small, times as long as broad, as long as its distance from the end of the snout, much shorter than the parietals. The
supraocular In scaled reptiles, supraocular scales are (enlarged) scales on the crown immediately above the eye.Mallow D, Ludwig D, Nilson G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Malabar, Florida: Krieger Publishing Company. ...
is as long as broad. There is one very small postocular. The temporals are arranged 1+1. There are seven
upper labials In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ( ...
, the third in contact with the prefrontal, the third and fourth (or third, fourth, and fifth) entering the eye. There are four lower labials in contact with the anterior
chin shield Chin shields or chinshields, genials scales on a snake are scales found on the underside of the snake's head towards the anterior and touching the lower labial scales. Chin shields to the front of the snake (towards the snout) are called anterio ...
. The two pairs of chin shields (anterior and posterior) are subequal in size. Boulenger GA (1896). ''Catalogue of the Snakes in the British Museum (Natural History). Volume III., containing the Colubridæ (Opisthoglyphæ and Proteroglyphæ), Amblycephalidæ, and Viperidæ.'' London: Trustees of the British Museum (Natural History). (Taylor and Francis, printers). xiv + 727 pp. + Plates I-XXV. (''Micrelaps vaillanti'', new combination, p. 249).


Reproduction

''M. vaillanti'' is
oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...
.


References


Further reading

* Mocquard F (1888). "''Sur une collection de reptiles et de batraciens rapportés des pays somalis et de Zanzibar par M'' nsieur ''G. Révoil'' ". ''Mémoires Publiés par la Société Philomatique à l'occasion du Centenaire de sa fondation 1788-1888, Paris''. pp. 109–134. (''Elaposchema vaillanti'', new species, p. 123 + Plate XII, figure 1). (in French). * Largen M, Spawls S (2010). ''Amphibians and Reptiles of Ethiopia and Eritrea''. Frankfurt am Main: Edition Chimaira / Serpents Tale. 694 pp. . (''Micrelaps vaillanti'', p. 586). *Spawls S, Howell K, Hinkel H, Menegon M (2018). ''Field Guide to East African Reptiles, Second Edition''. London: Bloomsbury Natural History. 624 pp. . (''Micrelaps vaillanti'', p. 459). {{Taxonbar, from=Q2712500 Atractaspididae Reptiles described in 1888