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''Mexcala meridiana'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Mexcala ''Mexcala'' is a genus of Salticidae, jumping spiders that was first described by George and Elizabeth Peckham in 1902. The name is probably derived from the Nahuatl ''mezcal''. Species it contains twenty-one species, found only in Africa, Yemen ...
'' that is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to South Africa, found in
Mpumalanga Mpumalanga () is one of the nine provinces of South Africa. The name means "East", or literally "The Place Where the Sun Rises" in the Nguni languages. Mpumalanga lies in eastern South Africa, bordering Eswatini and Mozambique. It shares bor ...
. The spider was first defined in 2009 by
Wanda Wesołowska Wanda Wesołowska (born 11 August 1950) is a Polish zoologist known for her work with jumping spiders. She has described more species of jumping spider than any contemporary writer, and is second only to Eugène Simon in the history of arachnolo ...
, one of over 500 that the
arachnologist Arachnology is the scientific study of arachnids, which comprise spiders and related invertebrates such as scorpions, pseudoscorpions, harvestmen, ticks, and mites. Those who study spiders and other arachnids are arachnologists. More narrowly ...
described during her career. It mimics ants and ant-like wasps, living alongside and preying upon them. Only the female has been identified. The spider is medium-sized, with a brown to blackish
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
typically long and a brownish
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
typically long. It is similar externally to other spiders in the genus, the most distinguishing feature being its copulatory organs.


Taxonomy

''Mexcala meridiana'' is a
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
that was first described by the Polish
arachnologist Arachnology is the scientific study of arachnids, which comprise spiders and related invertebrates such as scorpions, pseudoscorpions, harvestmen, ticks, and mites. Those who study spiders and other arachnids are arachnologists. More narrowly ...
Wanda Wesołowska Wanda Wesołowska (born 11 August 1950) is a Polish zoologist known for her work with jumping spiders. She has described more species of jumping spider than any contemporary writer, and is second only to Eugène Simon in the history of arachnolo ...
in 2009, one of over 500
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
she identified during her career. She allocated the species to the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Mexcala ''Mexcala'' is a genus of Salticidae, jumping spiders that was first described by George and Elizabeth Peckham in 1902. The name is probably derived from the Nahuatl ''mezcal''. Species it contains twenty-one species, found only in Africa, Yemen ...
'', first raised by George and Elizabeth Peckham in 1902 as part of a thorough revision of the genus. The genus was a member of the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Heliophaninae alongside '' Pseudicius'' and '' Cosmophasis'', which was absorbed into
Chrysillini Chrysillini is a tribe of jumping spider in the family Salticidae. In Maddison's 2015 revision of the family, the subfamily Heliophaninae was reclassified as a junior synonym of Chrysillini. Genera * '' Afraflacilla'' * '' Augustaea'' * '' ...
by
Wayne Maddison Wayne Paul Maddison (born 1958) is a Canadian evolutionary biologist, arachnologist, and biological illustrator. He is Canada Research Chair in Biodiversity and a professor at the departments of zoology and botany at the University of British ...
in 2015. The tribe is a member of the
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
Saltafresia within the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Salticoida Salticoida is an unranked clade of the jumping spider family Salticidae. It is the larger and more widespread of the two subdivisions of the "typical" jumping spiders (subfamily Salticinae), occurring effectively world-wide. Its sister clade is ...
. A year later, in 2016,
Jerzy Prószyński Jerzy Prószyński (born 1935 in Warsaw) is a Polish arachnologist specializing in systematics of jumping spiders (family Salticidae). He is a graduate of the University of Warsaw, a long-term employee of the Siedlce University of Natural Sc ...
allocated the genus to the Heliophanines group of genera, which was named after the genus '' Heliophanus''. The genera share characteristics, including having a rather uniform, mainly dark appearance. The species itself has a name that is derived from the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
word for southern and refers to it being found in Southern Africa.


Description

Like all ''Mexcala'' spiders, the species is slender and medium-sized. The female has a brown
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
covered in dense brown hairs that is typically long. The eye field is darker with black markings around the eyes. It has light brown mouthparts with yellow edges to the
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
. The
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
is typically long, brownish on top and darker underneath, with a pattern of two lighter streaks It has light brown legs. The
epigyne The epigyne or epigynum is the external genital structure of female spiders. As the epigyne varies greatly in form in different species, even in closely related ones, it often provides the most distinctive characteristic for recognizing species. ...
has a large shallow depression. The copulatory canals are separate to each other and lead to very simple internal copulatory organs. This differentiates the species from others in the genus. The male has not been described.


Behaviour

Like many jumping spiders, Wesołowska and Tamás Szűts noted that ''Mexcala'' spiders mimic ants. The species lives amongst various different species of ant and ant-like wasps that it mimics, and preys upon. Like other jumping spiders, it is a mainly diurnal hunter that uses its good eyesight to spot its prey. It attacks from the front and captures its prey behind the head. The chelicerae have short thick spines on them which may be used for digging holes to act as underground hiding places. It uses visual displays during courtship and transmits vibratory signals through silk to communicate to other spiders.


Distribution

''Mexcala'' spiders can be found across Africa and the Arabian peninsula. ''Mexcala meridiana'' is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to South Africa. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
was found near Sabie in
Mpumalanga Mpumalanga () is one of the nine provinces of South Africa. The name means "East", or literally "The Place Where the Sun Rises" in the Nguni languages. Mpumalanga lies in eastern South Africa, bordering Eswatini and Mozambique. It shares bor ...
in 1927.


References


Citations


Bibliography

* * * * * * * * {{Taxonbar, from=Q2144945 Endemic spiders of South Africa Salticidae Spiders described in 2009 Spiders of South Africa Taxa named by Wanda Wesołowska