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''Mexcala angolensis'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Mexcala ''Mexcala'' is a genus of Salticidae, jumping spiders that was first described by George and Elizabeth Peckham in 1902. The name is probably derived from the Nahuatl ''mezcal''. Species it contains twenty-one species, found only in Africa, Yemen ...
'' that is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to Angola, after which it is named. The spider was first defined in 2009 by
Wanda Wesołowska Wanda Wesołowska (born 11 August 1950) is a Polish zoologist known for her work with jumping spiders. She has described more species of jumping spider than any contemporary writer, and is second only to Eugène Simon in the history of arachnolo ...
, one of over 500 that the
arachnologist Arachnology is the scientific study of arachnids, which comprise spiders and related invertebrates such as scorpions, pseudoscorpions, harvestmen, ticks, and mites. Those who study spiders and other arachnids are arachnologists. More narrowly ...
described during her career. The spider is medium-sized, with a brownish
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
typically long and a light brown
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
typically long that has an indistinct dark streak that runs down the middle. It mimics ants and ant-like wasps, living alongside and preying upon them. The male can be distinguished from other ''Mexcala'' species by the small teeth at the end of its tibial apophysis, or spike, and the lack of a bump on its palpal bulb. The female has not been described.


Taxonomy

''Mexcala angolensis'' is a
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
that was first described by the Polish
arachnologist Arachnology is the scientific study of arachnids, which comprise spiders and related invertebrates such as scorpions, pseudoscorpions, harvestmen, ticks, and mites. Those who study spiders and other arachnids are arachnologists. More narrowly ...
Wanda Wesołowska Wanda Wesołowska (born 11 August 1950) is a Polish zoologist known for her work with jumping spiders. She has described more species of jumping spider than any contemporary writer, and is second only to Eugène Simon in the history of arachnolo ...
in 2009, one of over 500
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
she identified during her career. She allocated the species to the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Mexcala ''Mexcala'' is a genus of Salticidae, jumping spiders that was first described by George and Elizabeth Peckham in 1902. The name is probably derived from the Nahuatl ''mezcal''. Species it contains twenty-one species, found only in Africa, Yemen ...
'', first raised by George and Elizabeth Peckham in 1902 as part of a thorough revision of the genus. The genus was a member of the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Heliophaninae alongside '' Pseudicius'' and '' Cosmophasis'', which was absorbed into
Chrysillini Chrysillini is a tribe of jumping spider in the family Salticidae. In Maddison's 2015 revision of the family, the subfamily Heliophaninae was reclassified as a junior synonym of Chrysillini. Genera * '' Afraflacilla'' * '' Augustaea'' * '' ...
by
Wayne Maddison Wayne Paul Maddison (born 1958) is a Canadian evolutionary biologist, arachnologist, and biological illustrator. He is Canada Research Chair in Biodiversity and a professor at the departments of zoology and botany at the University of British ...
in 2015. The tribe is a member of the
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
Saltafresia within the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Salticoida Salticoida is an unranked clade of the jumping spider family Salticidae. It is the larger and more widespread of the two subdivisions of the "typical" jumping spiders (subfamily Salticinae), occurring effectively world-wide. Its sister clade is ...
. A year later, in 2016,
Jerzy Prószyński Jerzy Prószyński (born 1935 in Warsaw) is a Polish arachnologist specializing in systematics of jumping spiders (family Salticidae). He is a graduate of the University of Warsaw, a long-term employee of the Siedlce University of Natural Sc ...
allocated the genus to the Heliophanines group of genera, which was named after the genus '' Heliophanus''. The genera share characteristics, including having a rather uniform, mainly dark appearance. The species itself has a name that is derived from the country where it was first found.


Description

Like all ''Mexcala'' spiders, the species is slender and medium-sized. The male is covered with short brown hairs. a brownish
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
that is typically long. It has a short eye field. The
chelicerae The chelicerae () are the arthropod mouthparts, mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated ...
have a single tooth. The
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
is typically long and light brown with two lighter patches to the front and an indistinct dark streak down the middle. The underside has sets of yellowish dots forming four lines. The long thin legs are brown with black lines. The
pedipalp Pedipalps (commonly shortened to palps or palpi) are the secondary pair of forward appendages among Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to ...
s are brown with white scales on the tibia and cymbium and a rounded palpal bulb. There is a single tibial apophysis, or spike, with small teeth on its tip. This tip, along with the lack of a bump on the palpal bulb, help distinguish the species from others in the genus. The
embolus An embolus (; : emboli; from the Greek ἔμβολος "wedge", "plug") is an unattached mass that travels through the circulatory system, bloodstream and is capable of creating blockages. When an embolus Vascular occlusion, occludes a blood vess ...
is fixed to the tegulum. The female has not been described. Like many jumping spiders, Wesołowska and Tamás Szűts noted that ''Mexcala'' spiders mimic ants. Some are particularly similar to members of the ''
Camponotus Carpenter ants (''Camponotus'' spp.) are a genus of large ants (workers ) indigenous to many parts of the world. True carpenter ants build nests inside wood, consisting of galleries chewed out with their mandibles or jaws, preferably in dead, ...
'' genus. However, they mainly resemble Mutillidae, species of
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...
that have ant-like characteristics. The species particularly resembles the female, which is wingless, in its body proportions.


Behaviour

The spider is typical of the genus. It lives amongst various different species of ant and ant-like insect that it mimics, and preys upon. Like other jumping spiders, it is a mainly diurnal hunter that uses its good eyesight to spot its prey. It attacks from the front and captures its prey behind the head. The chelicerae have short thick spines on them that may be used for digging holes to act as underground hiding places. It uses visual displays during courtship and transmits vibratory signals through silk to communicate to other spiders.


Distribution and habitat

''Mexcala'' spiders can be found across Africa and the Arabian peninsula. ''Mexcala angolensis'' is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to Angola in Southern Africa. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
was found near Muconda in 1949.


References


Citations


Bibliography

* * * * * * * * * {{Taxonbar, from=Q2075724 Endemic fauna of Angola Fauna of Angola Salticidae Spiders of Africa Spiders described in 2009 Taxa named by Wanda Wesołowska