The Methylococcaceae are a family of

bacteria Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micr ...

that obtain their

carbon Carbon () is a chemical element; it has chemical symbol, symbol C and atomic number 6. It is nonmetallic and tetravalence, tetravalent—meaning that its atoms are able to form up to four covalent bonds due to its valence shell exhibiting 4 ...

and energy from

methane Methane ( , ) is a chemical compound with the chemical formula (one carbon atom bonded to four hydrogen atoms). It is a group-14 hydride, the simplest alkane, and the main constituent of natural gas. The abundance of methane on Earth makes ...

, called

methanotroph Methanotrophs (sometimes called methanophiles) are prokaryotes that metabolize methane as their source of carbon and chemical energy. They are bacteria or archaea, can grow aerobically or anaerobically, and require single-carbon compounds to ...

s.. They comprise the type I methanotrophs, in contrast to the

Methylocystaceae The Methylocystaceae are a family of bacteria that are capable of obtaining carbon and energy from methane. Such bacteria are called methanotrophs, and in particular the Methylocystaceae comprise the type II methanotrophs, which are structurally ...

or type II methanotrophs. They belong to

Gammaproteobacteria ''Gammaproteobacteria'' is a class of bacteria in the phylum ''Pseudomonadota'' (synonym ''Proteobacteria''). It contains about 250 genera, which makes it the most genus-rich taxon of the Prokaryotes. Several medically, ecologically, and scienti ...

, and are typically given their own

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

. The Methylococcaceae have internal membranes in the form of flattened discs, perpendicular to the

cell wall A cell wall is a structural layer that surrounds some Cell type, cell types, found immediately outside the cell membrane. It can be tough, flexible, and sometimes rigid. Primarily, it provides the cell with structural support, shape, protection, ...

. Methane is oxidized to give

formaldehyde Formaldehyde ( , ) (systematic name methanal) is an organic compound with the chemical formula and structure , more precisely . The compound is a pungent, colourless gas that polymerises spontaneously into paraformaldehyde. It is stored as ...

, which is fixed by a process called the ribulose monophosphate (RuMP) cycle. Here

is combined with sugar

ribulose Ribulose is a ketopentose — a monosaccharide containing five carbon atoms, and including a ketone functional group. It has chemical formula . Two enantiomers are possible, -ribulose (-erythro-pentulose) and -ribulose (-erythro-pentulose). ...

, producing hexulose. This, in turn, is broken down to produce

glyceraldehyde Glyceraldehyde (glyceral) is a triose monosaccharide with chemical formula C3 H6 O3. It is the simplest of all common aldoses. It is a sweet, colorless, crystalline solid that is an intermediate compound in carbohydrate metabolism. The word comes ...

, which is used to produce new ribulose and other organic compounds.

Catabolism Catabolism () is the set of metabolic pathways that breaks down molecules into smaller units that are either oxidized to release energy or used in other anabolic reactions. Catabolism breaks down large molecules (such as polysaccharides, lipid ...

does not involve a complete

citric acid cycle The citric acid cycle—also known as the Krebs cycle, Szent–Györgyi–Krebs cycle, or TCA cycle (tricarboxylic acid cycle)—is a series of chemical reaction, biochemical reactions that release the energy stored in nutrients through acetyl-Co ...

.George M. Garrity: ''

Bergey's Manual of Systematic Bacteriology ''Bergey's Manual of Systematic Bacteriology'' is the main resource for determining the identity of prokaryotic organisms, emphasizing bacterial species, using every characterizing aspect. The manual was published subsequent to ''Bergey's Manual ...

''. 2. Auflage. Springer, New York, 2005, Volume 2: ''The Proteobacteria, Part B: The Gammaproteobacteria'' Some species of the Methylococcaceae have formed with certain marine mussels endosymbiotic relationships.

References

External links

Methylococcaceae
J.P. Euzéby: List of Prokaryotic names with Standing in Nomenclature Gammaproteobacteria {{gammaproteobacteria-stub