Meteoraspis
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''Meteoraspis'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of ptychopariid
trilobite Trilobites (; meaning "three-lobed entities") are extinction, extinct marine arthropods that form the class (biology), class Trilobita. One of the earliest groups of arthropods to appear in the fossil record, trilobites were among the most succ ...
s of the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Tricrepicephalidae. The various species lived from 501 to 497 million years ago during the
Dresbachian The Dresbachian is a Maentwrogian regional stage of North America, lasting from 501 to 497 million years ago. It is part of the Upper Cambrian and is defined by four trilobite zones. It overlaps with the ICS-stages Guzhangian, Paibian and the ...
faunal stage In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by conven ...
of the late
Cambrian Period The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordovici ...
. Fossils of ''Meteoraspis'' are characteristic of Late Cambrian strata in North America, though they are found in Late Cambrian strata elsewhere in the world, such as ''M. nevensis'' from
Victoria Land Victoria Land is a region in eastern Antarctica which fronts the western side of the Ross Sea and the Ross Ice Shelf, extending southward from about 70°30'S to 78th parallel south, 78°00'S, and westward from the Ross Sea to the edge of the Ant ...
, Antarctica.


Description

The outline of the
exoskeleton An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. human skeleton, that ...
of ''Meteoraspis'' is an elongate, slightly inverted egg shaped though almost ovate, widest at the tip of the genal spines and 1.6× as long as wide. The headshield (or
cephalon Cephalon, Inc. was an American biopharmaceutical company co-founded in 1987 by pharmacologist Frank Baldino Jr., Frank Baldino, Jr., neuroscientist Michael Lewis, and organic chemist James C. Kauer—all three former scientists with the DuPont ...
) is highly vaulted. The well-defined central raised area (or
glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior ...
) is convex, elongate tapering, with a rounded front, and has only one furrow, crossing it near the back of the cephalon, defining the occipital ring. A node may be present on the occipital ring. The remaining parts of the cephalon, called fixed and free cheeks (or fixigenae and librigenae) are upsloping. The fracture lines (or sutures) that in moulting separate the librigenae from the fixigenae are divergent just in front of the eyes, becoming parallel near the border furrow and slightly convergent at the margin. From the back of the eyes the sutures follow a lazy S-curve, diverging first more outward and near the margin more backward, cutting the posterior margin within the inner bend of the spine (or
opisthoparian Trilobites (; meaning "three-lobed entities") are extinct marine arthropods that form the class Trilobita. One of the earliest groups of arthropods to appear in the fossil record, trilobites were among the most successful of all early animals, ...
sutures). Uniquely, the furrow parallel to the margin (or border furrow) has 2 not very noticeable pits in front of the glabella, and rarely a very faint 3rd median pit. There are no ridges that connect the eye to the glabella. The medium-sized eyes (over ⅓× the length of the glabella) are slightly behind the middle of the glabella. Genal angle ending in short spines extending backward about one thorax segment. The articulating middle part of the body (or
thorax The thorax (: thoraces or thoraxes) or chest is a part of the anatomy of mammals and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main di ...
) has 13 segments. The axis is convex, and less than ½× as wide as each of the so-called pleurae to its sides. Segments pointed sideways with a rounded front (a shape called falcate). The tailshield (or
pygidium The pygidium (: pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is compos ...
) is about ½× as wide as the cephalon, almost twice as wide as long, excluding the two flat, shark tooth shaped, widely spaced spines. The axis in the pygidium is 1¼× longer than wide, with almost parallel sides, almost reaching the rear margin, with 3 or 4 axial rings; 3 sets of interpleural grooves and pleural furrows ending at distance of the margin. There is no furrow that would define a border in the pygidium.


Comparison with ''Tricrepicephalus''

Species of the related genus '' Tricrepicephalus'' can be differentiated from those of ''Meteoraspis'' by having three equally prominent pits in the anterior border furrow, a much less vaulted cephalon, with spines reaching at least to the third thorax segment, 12 thorax segments and two long, tubular, curved pygidial spines that are reminiscent of the pincers of an
earwig Earwigs make up the insect order (biology), order Dermaptera. With about 2,000 species in 12 families, they are one of the smaller insect orders. Earwigs have characteristic cercus, cerci, a pair of forceps-like pincer (biology), pincers on ...
.


Reassigned species

Philippe Janvier Philippe Janvier is a French paleontology, paleontologist, specialising in Palaeozoic vertebrates, who currently works at the Muséum national d'Histoire naturelle, Museum National de l’Histoire Naturelle in Paris. He has written several books an ...
assigned ''Cephalaspis gigas'', an Early
Devonian The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ...
osteostracan The class Osteostraci (meaning "bony shells") is an extinct taxon of bony-armored jawless fish, termed "ostracoderms", that lived in what is now North America, Europe and Russia from the Middle Silurian to Late Devonian. Anatomically speaking, ...
agnathan with a tremendously wide headshield, to the newly erected genus ''Meteoraspis'' in 1981. This name however, was already used by Resser in 1935 for a trilobite. In 1987 the osteostracan genus was renamed to ''Parameteoraspis''. * ''M. caroli'' = '' Parameteoraspis caroli'' * ''M. gigas'' = '' Parameteoraspis gigas'' * ''M. lanternaria'' = '' Parameteoraspis laternaria'' * ''M. lata'' = '' Parameteoraspis lata'' * ''M. laticornis'' = '' Parameteoraspis laticornis'' * ''M. menoides'' = '' Parameteoraspis menoides'' * ''M. oberon'' = '' Parameteoraspis oberon'' * ''M. oblonga'' = '' Parameteoraspis oblonga''


Distribution

* ''M. etheridgei'' is known from the Cambrian of Australia (late Mindyallan, Dolodrook River Limestone Formation, Garvey Gully, Dolodrook River, East Victoria, 37.6° S, 146.6° E). ''cited in'' * ''M. metra'' has been excavated from the Upper Cambrian of the United States (Dresbachian, ''Coosella'' zone, Riley Formation, Central Texas, 30.3° N, 97.7° W; ''cited in'' and ''Crepicephalus'' trilobite zone, lower Member of the Deadwood Formation, Whitewood Creek in Lead County, South Dakota, 44.3° N, 103.8° W ''cited in'' ).


Ecology

''M. etheridgei'' occurs together with ''Eugonocare tessellatum'', ''Corynexochus plumula'', ''Mindycrusta oepiki'', ''Innitagnostus inexpectans'', ''Peratagnostus'', and ''Pseudagnostus idalis'' (all trilobites), ''Acrothele'', ''Orbithele'', ''Quadrisonia'', ''Anabolotreta tegula'', ''Anabolotreta sp.'', ''Dactylotreta redunca'', ''Dactylotreta sp.'', ''Linnarssonia'', ''Neotreta'', ''Picnotreta sp.'', ''Picnotreta debilis'', ''Stilpnotreta magna'', ''Treptotreta sp.'', ''Treptotreta jucunda'', ''Dictyonina'', ''Micromitra modesta'' and ''Micromitra sp.'' (all brachiopods).


References

{{Taxonbar, from=Q6823448 Ptychopariida genera Ptychopariida Cambrian trilobites