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The metatarsal bones or metatarsus (: metatarsi) are a group of five
long bone The long bones are those that are longer than they are wide. They are one of five types of bones: long, short, flat, irregular and sesamoid. Long bones, especially the femur and tibia, are subjected to most of the load during daily activities ...
s in the
midfoot The foot (: feet) is an anatomical structure found in many vertebrates. It is the terminal portion of a limb which bears weight and allows locomotion. In many animals with feet, the foot is an organ at the terminal part of the leg made up of ...
, located between the tarsal bones (which form the
heel The heel is the prominence at the posterior end of the foot. It is based on the projection of one bone, the calcaneus or heel bone, behind the articulation of the bones of the lower leg. Structure To distribute the compressive forces exerted ...
and the
ankle The ankle, the talocrural region or the jumping bone (informal) is the area where the foot and the leg meet. The ankle includes three joints: the ankle joint proper or talocrural joint, the subtalar joint, and the inferior tibiofibular joint. The ...
) and the
phalanges The phalanges (: phalanx ) are digit (anatomy), digital bones in the hands and foot, feet of most vertebrates. In primates, the Thumb, thumbs and Hallux, big toes have two phalanges while the other Digit (anatomy), digits have three phalanges. ...
(
toe Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being ''digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being ''plantigrade''; ...
s). Lacking individual names, the metatarsal bones are numbered from the medial side (the side of the
great toe Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being ''digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being ''plantigrade''; ...
): the first,
second The second (symbol: s) is a unit of time derived from the division of the day first into 24 hours, then to 60 minutes, and finally to 60 seconds each (24 × 60 × 60 = 86400). The current and formal definition in the International System of U ...
,
third Third or 3rd may refer to: Numbers * 3rd, the ordinal form of the cardinal number 3 * , a fraction of one third * 1⁄60 of a ''second'', i.e., the third in a series of fractional parts in a sexagesimal number system Places * 3rd Street (di ...
, fourth, and fifth metatarsal (often depicted with
Roman numerals Roman numerals are a numeral system that originated in ancient Rome and remained the usual way of writing numbers throughout Europe well into the Late Middle Ages. Numbers are written with combinations of letters from the Latin alphabet, eac ...
). The metatarsals are analogous to the
metacarpal In human anatomy, the metacarpal bones or metacarpus, also known as the "palm bones", are the appendicular bones that form the intermediate part of the hand between the phalanges (fingers) and the carpal bones ( wrist bones), which articulate ...
bones of the
hand A hand is a prehensile, multi-fingered appendage located at the end of the forearm or forelimb of primates such as humans, chimpanzees, monkeys, and lemurs. A few other vertebrates such as the Koala#Characteristics, koala (which has two thumb#O ...
. The lengths of the metatarsal bones in humans are, in descending order, second, third, fourth, fifth, and first. A bovine hind leg has two metatarsals.


Structure

The five metatarsals are
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
convex long bones consisting of a shaft or body, a base (
proximal Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...
ly), and a head (
distal Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provi ...
ly).Platzer 2004, p. 220 The body is prismoid in form, tapers gradually from the tarsal to the phalangeal extremity, and is curved longitudinally, so as to be concave below, slightly convex above. The base or posterior extremity is
wedge A wedge is a triangle, triangular shaped tool, a portable inclined plane, and one of the six simple machines. It can be used to separate two objects or portions of an object, lift up an object, or hold an object in place. It functions by conver ...
-shaped, articulating proximally with the tarsal bones, and by its sides with the contiguous metatarsal bones: its dorsal and plantar surfaces are rough for the attachment of
ligament A ligament is a type of fibrous connective tissue in the body that connects bones to other bones. It also connects flight feathers to bones, in dinosaurs and birds. All 30,000 species of amniotes (land animals with internal bones) have liga ...
s. The head or distal extremity presents a convex articular surface, oblong from above downward, and extending farther backward below than above. Its sides are flattened, and on each is a depression, surmounted by a
tubercle In anatomy, a tubercle (literally 'small tuber', Latin for 'lump') is any round nodule, small eminence, or warty outgrowth found on external or internal organs of a plant or an animal. In plants A tubercle is generally a wart-like projectio ...
, for ligamentous attachment. Its
plantar Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provi ...
surface is grooved antero-posteriorly for the passage of the flexor
tendon A tendon or sinew is a tough band of fibrous connective tissue, dense fibrous connective tissue that connects skeletal muscle, muscle to bone. It sends the mechanical forces of muscle contraction to the skeletal system, while withstanding tensi ...
s, and marked on either side by an articular eminence continuous with the terminal articular surface.Gray's 1918, ''6d. 2. The Metatarsus'' During growth, the
growth plate The epiphyseal plate, epiphysial plate, physis, or growth plate is a hyaline cartilage plate in the metaphysis at each end of a long bone. It is the part of a long bone where new bone growth takes place; that is, the whole bone is alive, with ma ...
s are located distally on the metatarsals, except on the first metatarsal where it is located proximally. Yet it is quite common to have an accessory growth plate on the distal first metatarsal.


Articulations

The base of each metatarsal bone articulates with one or more of the tarsal bones at the tarsometatarsal joints, and the head with one of the first row of phalanges at the
metatarsophalangeal joint The metatarsophalangeal joints (MTP joints) are the joints between the metatarsal bones of the foot and the proximal bones (proximal phalanges) of the toes. They are analogous to the knuckles of the hand, and are consequently known as toe knuckles ...
s. Their bases also articulate with each other at the intermetatarsal joints * The ''first'' metatarsal articulates with the medial cuneiform, and to a small extent to the intermediate cuneiform.Platzer 2004, p. 218 * the ''second'' with all three cuneiforms. * the ''third'' with the lateral cuneiform. * the ''fourth'' with the lateral cuneiform and the
cuboid In geometry, a cuboid is a hexahedron with quadrilateral faces, meaning it is a polyhedron with six Face (geometry), faces; it has eight Vertex (geometry), vertices and twelve Edge (geometry), edges. A ''rectangular cuboid'' (sometimes also calle ...
. * The ''fifth'' with the cuboid.


Muscle attachments


Clinical significance


Injuries

The metatarsal bones are often broken by
association football Association football, more commonly known as football or soccer, is a team sport played between two teams of 11 Football player, players who almost exclusively use their feet to propel a Ball (association football), ball around a rectangular f ...
(soccer) players. These and other recent cases have been attributed to the lightweight design of modern
football boot Football boots, also referred to as cleats or soccer shoes in North America, are specialized footwear designed for use in association football. Those designed for grass Association football pitch, pitches have cleat (shoe), studs on the outsole ...
s, which provide less protection to the foot. In 2010 some football players began testing a new sock that incorporated a rubber silicone pad over the foot to provide protection to the top of the foot. Stress fractures are thought to account for 16% of injuries related to sports participation, and the metatarsals are the bones most often involved. These fractures are sometimes called march fractures, based on their traditional association with military recruits after long marches. The second and third metatarsals are fixed while walking, thus these metatarsals are common sites of injury. The fifth metatarsal may be fractured if the foot is oversupinated during locomotion. Protection from injuries can be given by the use of safety footwear which can use built-in or removable metatarsal guards.


Additional images

Image:Footx.jpg,
X-ray An X-ray (also known in many languages as Röntgen radiation) is a form of high-energy electromagnetic radiation with a wavelength shorter than those of ultraviolet rays and longer than those of gamma rays. Roughly, X-rays have a wavelength ran ...
of foot. Image:Gray290 - Mratatarsus.png, Skeleton of left foot. Medial aspect. Image:Gray360.png, Oblique section of left intertarsal and tarsometatarsal articulations, showing the synovial cavities. File:Slide2xzxzx.JPG, Ankle and tarsometarsal joints, showing bones of foot. Deep dissection. File:Nitti MetatarsalGuard.jpg, alt=Nitti Safety Footwear with removable metatarsal guard., Safety footwear with removable metatarsal guard.


See also

*
Arches of the foot The arches of the foot, formed by the tarsal and metatarsal bones, strengthened by ligaments and tendons, allow the foot to support the weight of the body in the erect posture with the least weight. They are categorized as longitudinal and t ...
*
Ball (foot) The ball of the foot is the padded portion of the sole between the toes and the arch, underneath the heads of the metatarsal bones. In comparative foot morphology, the ball is most analogous to the metacarpal (forepaw) or metatarsal (hindpa ...
* Bone terminology *
Terms for anatomical location Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Thi ...
*
Jones fracture A Jones fracture is a broken bone in a specific part of the fifth metatarsal of the foot between the base and middle part that is known for its high rate of delayed healing or nonunion. It results in pain near the midportion of the foot on ...
* Lisfranc injury * Morton's toe


Notes


References

* *


External links

* {{Authority control Bones of the lower limb Bones of the foot