''Mesolimulus'' is an extinct
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
horseshoe crab
Horseshoe crabs are arthropods of the family Limulidae and the only surviving xiphosurans. Despite their name, they are not true crabs or even crustaceans; they are chelicerates, more closely related to arachnids like spiders, ticks, and scor ...
. The best known examples are found in
Solnhofen limestone
The Solnhofen Limestone or Solnhofen Plattenkalk, formally known as the Altmühltal Formation, is a Jurassic Konservat-Lagerstätte that preserves a rare assemblage of fossilized organisms, including highly detailed imprints of soft bodied organi ...
near
Solnhofen
Solnhofen is a municipality in the district of Weißenburg-Gunzenhausen in the region of Middle Franconia in the ' of Bavaria in Germany. It is in the Altmühl valley.
The local area is famous in geology and palaeontology for Solnhofen lime ...
,
Bavaria
Bavaria, officially the Free State of Bavaria, is a States of Germany, state in the southeast of Germany. With an area of , it is the list of German states by area, largest German state by land area, comprising approximately 1/5 of the total l ...
,
Germany
Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
. Originally assigned to the living genus ''
Limulus'', they are related to and look virtually identical to modern horseshoe crabs. Other species assigned to ''Mesolimulus'' have been recorded spanning over 140 million years from the
Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is di ...
to
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
from
England
England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
,
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
,
Siberia
Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
and
Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
.
''Kouphichnium''
The unusual
trackway
Historic roads (or historic trails in the US and Canada) are paths or routes that have historical importance due to their use over a period of time. Examples exist from prehistoric times until the early 20th century. They include ancient track ...
s left by these animals at Solnhofen were originally thought to have been made by
birds
Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
or
pterosaurs
Pterosaurs are an extinct clade of flying reptiles in the Order (biology), order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosau ...
, since they included a cross-shaped marking comparable with the impression of a typical bird foot. Eventually they were correctly identified as arthropod footprints and named ''Kouphichnium walchi'', as proved by some remarkable 'death traces' showing a ''Mesolimulus'' circling round on itself before dying. The last leg of more advanced horseshoe crabs is modified into a so-called 'pusher' which consists of four plates at the tip which push against the soft sediment rather like a snow-shoe. This was what left the unusual bird-like footprints.
Horseshoe crabs in general date to the
Ordovician Period
The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era, and the second of twelve periods of the Phanerozoic Eon. The Ordovician spans 41.6 million years from the end of the Cambrian Period Ma (mill ...
, more than 440 million years ago, and late
Paleozoic
The Paleozoic ( , , ; or Palaeozoic) Era is the first of three Era (geology), geological eras of the Phanerozoic Eon. Beginning 538.8 million years ago (Ma), it succeeds the Neoproterozoic (the last era of the Proterozoic Eon) and ends 251.9 Ma a ...
''
Euproops'' fossils indicate that they have changed little over the last 300 million years. Fossils preserved in Solnhofen limestone are unusual because soft body parts and skeletons are clearly represented.
Species
After Bicknell ''et al.'' (2021):
* ''Mesolimulus sibiricus'' Ponomarenko, 1985,
Middle Jurassic
The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
(
Bathonian
In the geologic timescale the Bathonian is an age (geology), age and stage (stratigraphy), stage of the Middle Jurassic. It lasted from approximately 168.2 ±1.2 annum, Ma to around 165.3 ±1.1 Ma (million years ago). The Bathonian Age succeeds ...
)
Talynzhansk Formation, Russia
* ''Mesolimulus tafraoutensis'' Lamsdell et al., 2020
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(upper
Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
–lower
Turonian
The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
)
Gara Sbaa Lagerstatte, Morocco
* ''Mesolimulus walchi'' (Desmarest, 1822)
Late Jurassic
The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
(
Kimmeridgian
In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
–Early
Tithonian
In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
)
Solnhofen Limestone
The Solnhofen Limestone or Solnhofen Plattenkalk, formally known as the Altmühltal Formation, is a Jurassic Konservat-Lagerstätte that preserves a rare assemblage of fossilized organisms, including highly detailed imprints of soft bodied organi ...
, Germany
* ''Mesolimulus woodwardi'' (Watson, 1909) Middle Jurassic (
Aalenian
The Aalenian () is a subdivision of the Middle Jurassic Epoch/Series of the geologic timescale that extends from about 174.7 ±0.8 Ma to about 170.9 ±0.8 Ma (million years ago). It was preceded by the Toarcian and succeeded by the Bajocian.
St ...
)
Northampton Sand Formation
The Northampton Sand Formation, sometimes called the Northamptonshire Sand, is a Middle Jurassic geological formation which is placed within the Inferior Oolite Group. It was formerly worked extensively in Northamptonshire for its ironstone.
...
, England
* ''Mesolimulus crespelli'' Vıa Boada, 1987
Middle Triassic
In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is di ...
(
Ladinian
The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between Ma and ~237 Ma (million years ago). The Ladinian was preceded by the Anisian and succeeded by the Carnian (part of the Upper or Late Triassic ...
)
Alcover Limestone Formation, Spain
References
*
*
Solnhofen fauna
Xiphosura
Late Jurassic arthropods
{{paleo-arthropod-stub