The cephalic index or cranial index is a number obtained by taking the maximum width ( biparietal diameter or BPD, side to side) of the head of an

organism An organism is any life, living thing that functions as an individual. Such a definition raises more problems than it solves, not least because the concept of an individual is also difficult. Many criteria, few of them widely accepted, have be ...

, multiplying it by 100 and then dividing it by their maximum length ( occipitofrontal diameter or OFD, front to back). The index was once used to categorize human beings in the first half of the 20th century, but today it is used to categorize dogs and cats.

Historic use in anthropology

Early anthropology

The cephalic index was used by anthropologists in the early 20th century as a tool to categorize human populations. It was used to describe an individual's appearance and for estimating the age of

fetus A fetus or foetus (; : fetuses, foetuses, rarely feti or foeti) is the unborn offspring of a viviparous animal that develops from an embryo. Following the embryonic development, embryonic stage, the fetal stage of development takes place. Pren ...

es for legal and obstetrical reasons. The cephalic index was defined by Swedish professor of

anatomy Anatomy () is the branch of morphology concerned with the study of the internal structure of organisms and their parts. Anatomy is a branch of natural science that deals with the structural organization of living things. It is an old scien ...

Anders Retzius (1796–1860) and first used in

physical anthropology Biological anthropology, also known as physical anthropology, is a natural science discipline concerned with the biological and behavioral aspects of human beings, their extinct hominin ancestors, and related non-human primates, particularly from ...

to classify ancient human remains found in Europe. The theory became closely associated with the development of racial anthropology in the 19th and early 20th centuries, when historians attempted to use ancient remains to model population movements in terms of racial categories. American anthropologist Carleton S. Coon also used the index in the 1960s, by which time it had been largely discredited. PSM V59 D404 Cephalic indexes of skull shapes

PSM V59 D404 Cephalic indexes of skull shapes

In the cephalic index model, human beings were characterized by having either a dolichocephalic (long-headed), mesaticephalic (moderate-headed), or brachycephalic (short-headed) cephalic index or cranial index. Otmaa1917 1-1-

Indices

Cephalic indices are grouped as in the following table: Technically, the measured factors are defined as the maximum width of the bones that surround the head above the supramastoid crest (behind the cheekbones), and the maximum length from the most easily noticed part of the

glabella The glabella, in humans, is the area of skin between the eyebrows and above the nose. The term also refers to the underlying bone that is slightly depressed, and joins the two brow ridges. It is a cephalometric landmark that is just superior ...

(between the eyebrows) to the most easily noticed point on the back part of the head.

Controversy

The usefulness of the cephalic index was questioned by the Italian anthropologist

Giuseppe Sergi Giuseppe Sergi (March 20, 1841 – October 17, 1936) was an Italian anthropologist of the early twentieth century, best known for his opposition to Nordicism in his books on the racial identity of Mediterranean peoples. He rejected existing ra ...

, who argued that cranial morphology provided a better means to model racial ancestry. Also,

Franz Boas Franz Uri Boas (July 9, 1858 – December 21, 1942) was a German-American anthropologist and ethnomusicologist. He was a pioneer of modern anthropology who has been called the "Father of American Anthropology". His work is associated with the mov ...

studied the children of immigrants to the United States in 1910 to 1912, noting that the children's cephalic index differed significantly from their parents', implying that local environmental conditions had a significant effect on the development of head shape. Boas argued that if craniofacial features were so malleable in a single generation, then the cephalic index was of little use for defining race and mapping ancestral populations. Scholars such as Earnest Hooton continued to argue that both environment and heredity were involved. Boas did not himself claim it was totally plastic. In 2002, a paper by Sparks and Jantz re-evaluated some of Boas's original data using new statistical techniques and concluded that there was a "relatively high genetic component" of head shape. Ralph Holloway of Columbia University argues that the new research raises questions about whether the variations in skull shape have "adaptive meaning and whether, in fact, normalizing selection might be at work on the trait, where both extremes, hyperdolichocephaly and hyperbrachycephaly, are at a slight selective disadvantage." In 2003, anthropologists Clarence C. Gravlee, H. Russell Bernard, and William R. Leonard reanalyzed Boas's data and concluded that most of Boas's original findings were correct. Moreover, they applied new statistical, computer-assisted methods to Boas's data and discovered more evidence for cranial plasticity. In a later publication, Gravlee, Bernard and Leonard reviewed Sparks's and Jantz's analysis. They argue that Sparks and Jantz misrepresented Boas's claims, and that Sparks's and Jantz's data support Boas. For example, they point out that Sparks and Jantz look at changes in cranial size in relation to how long an individual has been in the United States in order to test the influence of the environment. Boas, however, looked at changes in cranial size in relation to how long the mother had been in the United States. They argue that Boas's method is more useful, because the prenatal environment is a crucial developmental factor. Jantz and Sparks responded to Gravlee et al., reiterating that Boas' findings lacked biological meaning, and that the interpretation of Boas' results common in the literature was biologically inaccurate. In a later study, the same authors concluded that the effects Boas observed were likely the result of population-specific environmental effects such as changes in cultural practices for cradling infants, rather than the effects of a general "American environment" which caused populations in America to converge to a common cranial type, as Boas had suggested.

Vertical cephalic index

The vertical cephalic index, also known as the length-height index, was a less-commonly measured head ratio. In the vertical cephalic index model, humans beings were characterized by having either a chamaecranic (low-skulled), orthocranic (medium high-skulled), or hypsicranic (high-skulled) cephalic index or cranial index.

Medicine

The cephalic index is also used in medicine, especially in the planning and effectiveness analysis of cranial deformity corrections. The index is a useful tool in assessing the morphology of cranial deformities in clinical settings. The index is used while looking at the fetal head shape, and can change in certain situations (ex. breech presentation, ruptured membranes,

twin pregnancy Twins are two offspring produced by the same pregnancy.MedicineNet > Definition of Twin Last Editorial Review: 19 June 2000 Twins can be either ''monozygotic'' ('identical'), meaning that they develop from one zygote, which splits and forms two e ...

). It can be measured for prenatal infants in the womb using obstetric ultrasound, and a value of 78.3% ± 8% (±2

standard deviations In statistics, the standard deviation is a measure of the amount of variation of the values of a variable about its mean. A low standard deviation indicates that the values tend to be close to the mean (also called the expected value) of the ...

) is considered normal.

Modern use in animal breeding

The cephalic index is used in the categorisation of animals, especially breeds of dogs and cats.

Brachycephalic animals

A brachycephalic skull is relatively broad and short (typically with the breadth at least 80% of the length). Dog breeds such as the

pug The Pug is a breed of dog with the physically distinctive features of a wrinkly, short-muzzled face, and curled tail. An ancient breed, with roots dating back to 400 B.C., they have a fine, glossy coat that comes in a variety of colors, most ...

are sometimes classified as "extreme brachycephalic". Because of health issues brachycephaly is regarded in some countries as "qualzucht", which literally translates to "torture breeding" as it often leads to

brachycephalic airway obstructive syndrome Brachycephalic obstructive airway syndrome (BOAS), also known as brachycephalic airway obstructive syndrome (BAOS), brachycephalic airway syndrome (BAS), and brachycephalic syndrome (BS), is a pathological condition affecting brachycephalic, sho ...

List of brachycephalic dogs

* Affenpinscher *

American Bulldog The American bulldog is a large, muscular dog breed, breed of mastiff-type dog. Their ancestors were brought to the British North American colonies where they worked on small farms and ranches. History Dog breeds defined to any standard onl ...

* American Bully * Boston Terrier * Boxer * Brussels Griffon *

Bulldog The Bulldog is a British breed of dog of mastiff type. It may also be known as the English Bulldog or British Bulldog. It is a stocky, muscular dog of medium size, with a large head, thick folds of skin around the face and shoulders and a rel ...

* Bullmastiff * Cane Corso *

Cavalier King Charles Spaniel The Cavalier King Charles Spaniel (CKCS) is a British list of dog breeds, breed of toy dog of spaniel type. Four colours are recognised: Blenheim (chestnut and white), Tricolor (dog), tricolour (black/white/tan), black and tan, and ruby; the coa ...

* Apple-headed Chihuahua *

Chow Chow The Chow Chow is a spitz-type of dog breed originally from Northern China. The Chow Chow is a sturdily built dog, square in profile, with a broad skull and small, triangular, erect ears with rounded tips. The breed is known for a very dense do ...

Dogo Argentino The Dogo Argentino (plural Dogos Argentinos) is an Argentine list of dog breeds, breed of large dog of mastiff type. It was selective breeding, bred in the early twentieth century in Córdoba, Argentina, Córdoba in central Argentina, primarily ...

* Dogue de Bordeaux *

English Mastiff The English Mastiff, or simply the Mastiff, is a British list of dog breeds, breed of very large dog of mastiff type. It is likely descended from the ancient Alaunt and Pugnaces Britanniae, with a significant input from the Alpine Mastiff in th ...

* English Bulldog * Fila Brasileiro * French Bulldog * Japanese Chin *

King Charles Spaniel The King Charles Spaniel (also known as the English Toy Spaniel) is a small dog breed of the spaniel type. In 1903, The Kennel Club combined four separate toy spaniel breeds under this single title. The other varieties merged into this breed wer ...

* Lhasa Apso * Lowchen *

Neapolitan Mastiff The Neapolitan Mastiff or is an Italian breed of large dog of mastiff type. It was recognised as a breed by the Ente Nazionale della Cinofilia Italiana in 1949, and accepted by the Fédération Cynologique Internationale in 1956. It descends ...

Newfoundland Newfoundland and Labrador is the easternmost province of Canada, in the country's Atlantic region. The province comprises the island of Newfoundland and the continental region of Labrador, having a total size of . As of 2025 the population ...

* Olde English Bulldogge *

Pekingese The Pekingese (also spelled Pekinese) is a dog breed, breed of toy dog, originating in China. The breed was favored by royalty of the Chinese sovereign, Chinese Imperial court as a companion dog, and its name refers to the city of Beijing (Peki ...

* Perro de Presa Canario *

Pit bull Pit bull is an umbrella term for several Dog type, types of dog believed to have descended from bull and terriers. In the United States, the term is usually considered to include the American Pit Bull Terrier, American Staffordshire Terrier, A ...

Pug The Pug is a breed of dog with the physically distinctive features of a wrinkly, short-muzzled face, and curled tail. An ancient breed, with roots dating back to 400 B.C., they have a fine, glossy coat that comes in a variety of colors, most ...

* Pyrenean Mastiff * Shar-Pei * Shih Tzu * Tibetan Spaniel * Tosa

List of brachycephalic cats

Nipsu-Nappulan Non-Stop Exotic shorthair

British Shorthair The British Shorthair is the Purebred, pedigree version of the traditional British Domestic cat (landrace), domestic cat, with a distinctively stocky body, thick Fur, coat, and broad face. The most familiar colour variant is the "British Blue", wi ...

Burmese cat The Burmese cat (, , or , or , meaning copper colour) is a cat breed, breed of domestic cat, originating in Myanmar, Burma, believed to have its roots near the Myanmar–Thailand border, Thai–Burma border and developed in the United Sta ...

Exotic Shorthair The Exotic Shorthair is a breed of cat developed as a short-haired version of the Persian. The Exotic is similar to the Persian in appearance with the exception of the short dense coat. History In the late 1950s, the Persian was used as an ...

Himalayan cat The Himalayan (short for Himalayan Persian, or Colourpoint Persian as it is commonly referred to in Europe), is a breed or sub-breed of long-haired cat similar in type to the Persian cat, Persian, with the exception of its blue eyes and its poin ...

Persian cat The Persian cat, also known as the Persian Longhair, is a long-haired breed of cat characterised by a round face and short muzzle. The first documented ancestors of Persian cats might have been imported into Italy from Greater Khorasan, Khoras ...

Scottish Fold The Scottish Fold is a distinctive List of cat breeds, breed of Cat, domestic cat characterised by a natural Dominance (genetics), dominant gene mutation associated with osteochondrodysplasia. This genetic anomaly affects cartilage throughout t ...

White tiger The white tiger (ashy tiger) is a leucistic morph of the tiger, typically the Bengal tiger. It is occasionally reported in the Indian wilderness. It has the typical black stripes of a tiger, but its coat is otherwise white or near-white, and ...

Cheetah The cheetah (''Acinonyx jubatus'') is a large Felidae, cat and the Fastest animals, fastest land animal. It has a tawny to creamy white or pale buff fur that is marked with evenly spaced, solid black spots. The head is small and rounded, wit ...

Leopard The leopard (''Panthera pardus'') is one of the five extant cat species in the genus ''Panthera''. It has a pale yellowish to dark golden fur with dark spots grouped in rosettes. Its body is slender and muscular reaching a length of with a ...

Jaguar The jaguar (''Panthera onca'') is a large felidae, cat species and the only extant taxon, living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat spe ...

Snow Leopard The snow leopard (''Panthera uncia'') is a species of large cat in the genus ''Panthera'' of the family Felidae. The species is native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because ...

Mountain Lion The cougar (''Puma concolor'') (, ''Help:Pronunciation respelling key, KOO-gər''), also called puma, mountain lion, catamount and panther is a large small cat native to the Americas. It inhabits North America, North, Central America, Cent ...

List of brachycephalic pigs

* Middle White * Neijiang

List of brachycephalic rabbits

* Lionhead rabbit *

Lop rabbit Lop rabbit or lop-eared rabbit refers to any rabbit with ears that droop, as opposed to being carried erect. A number of rabbit breeds (listed below) are characterized by such lop ears. Abnormalities in the skull of a half-lop rabbit were studied ...

* Netherland Dwarf rabbit * Dwarf Papillon rabbit * Dwarf Hotot rabbit * Jersey Wooly rabbit * American Fuzzylop rabbit

Other

Elephant Elephants are the largest living land animals. Three living species are currently recognised: the African bush elephant ('' Loxodonta africana''), the African forest elephant (''L. cyclotis''), and the Asian elephant ('' Elephas maximus ...

Giant panda The giant panda (''Ailuropoda melanoleuca''), also known as the panda bear or simply panda, is a bear species endemic to China. It is characterised by its white animal coat, coat with black patches around the eyes, ears, legs and shoulders. ...

* Ross seal * Spectacled bear *

Walrus The walrus (''Odobenus rosmarus'') is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobeni ...

Wombat Wombats are short-legged, muscular quadrupedal marsupials of the family Vombatidae that are native to Australia. Living species are about in length with small, stubby tails and weigh between . They are adaptable and habitat tolerant, and are ...

* Australian Lowline * brachycephalic

Arabian horse The Arabian or Arab horse ( , DIN 31635, DMG ''al-ḥiṣān al-ʿarabī'') is a horse breed, breed of horse with historic roots on the Arabian Peninsula. With a distinctive head shape and high tail carriage, the Arabian is one of the most easi ...

* Niata cattle *

Tortoise Tortoises ( ) are reptiles of the family Testudinidae of the order Testudines (Latin for "tortoise"). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like o ...

Raccoon dog ''Nyctereutes'' (Greek: ''nyx, nykt-'' "night" + ''ereutēs'' "wanderer") is a genus of canid which includes only two extant species, both known as raccoon dogs: the common raccoon dog (''Nyctereutes procyonoides'') and the Japanese raccoon do ...

European bison The European bison (: bison) (''Bison bonasus'') or the European wood bison, also known as the wisent ( or ), the zubr (), or sometimes colloquially as the European buffalo, is a European species of bison. It is one of two extant species of bi ...

Otter Otters are carnivorous mammals in the subfamily Lutrinae. The 13 extant otter species are all semiaquatic, aquatic, or marine. Lutrinae is a branch of the Mustelidae family, which includes weasels, badgers, mink, and wolverines, among ...

Sloth Sloths are a Neotropical realm, Neotropical group of xenarthran mammals constituting the suborder Folivora, including the extant Arboreal locomotion, arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of move ...

Mesaticephalic animals

A mesaticephalic skull is of intermediate length and width. Mesaticephalic skulls are not markedly brachycephalic or dolichocephalic. When dealing with animals, especially dogs, the more appropriate and commonly used term is not "mesocephalic", but rather "mesaticephalic", which is a ratio of head to nasal cavity. The breeds below exemplify this category.

List of mesaticephalic canines

African Wild Dog The African wild dog (''Lycaon pictus''), also called painted dog and Cape hunting dog, is a wild canine native to sub-Saharan Africa. It is the largest wild canine in Africa, and the only extant member of the genus '' Lycaon'', which is disti ...

* Alaskan Malamute * almost all

spaniel A spaniel is a Dog type, type of gun dog. Spaniels were especially bred to flush game out of denser brush. By the late 17th century, spaniels had been specialized into water and land breeds. The extinct English Water Spaniel was used to retriev ...

s * almost all

spitz A spitz (; , in reference to the pointed muzzle) is a Dog type, type of domestic dog consisting of between 50 and 70 breeds depending on classification. There is no precise definition of 'spitz' but typically most spitz breeds have pricked ears ...

, except for the Chow Chow * American Eskimo Dog * American Foxhound * Appenzeller Sennenhund *

Australian Cattle Dog The Australian Cattle Dog, or simply Cattle Dog, is a breed of herding dog developed in Australia for droving cattle over long distances across rough terrain. This breed is a medium-sized, short-Coat (dog), coated dog that occurs in two main ...

Australian Shepherd The Australian Shepherd is a breed of herding dog from the United States. The name of the breed is technically a misnomer, as it was developed in California in the 19th century. It is believed to have its origins in sheepdog breeds from northwe ...

* Basenji *

Beagle The Beagle is a small breed of scent hound, similar in appearance to the much larger foxhound. The beagle was developed primarily for hunting rabbit or hare, known as beagling. Possessing a great sense of smell and superior tracking inst ...

Bearded Collie The Bearded Collie or Beardie is a British breed of herding dog of collie type. It was formerly used primarily by Scottish shepherds, but now commonly kept as a family companion. Weights are usually in the range , while height at the withers ...

Beauceron The Beauceron () is a herding dog breed originating from the plains of Central France. The Beauceron is also known as Berger de Beauce (, sheepdog from Beauce) or Bas Rouge (, red-stockings). Description Appearance This breed stands in height ...

* Belgian Malinois * Belgian Sheepdog * Bernese Mountain Dog * Bichon Frisé *

Black and Tan Coonhound The Black and Tan Coonhound is a breed of hunting dog. Developed in the United States from crosses between the Bloodhound and the Black and Tan Virginia Foxhound, this scent hound runs its game entirely by scent and is used primarily for racc ...

Border Collie The Border Collie is a British list of dog breeds, breed of herding dog of the collie type of medium size. It originates in the region of the Anglo-Scottish border, and descends from the traditional Sheep dog, sheepdogs once found all over the ...

Cardigan Welsh Corgi The Cardigan Welsh corgi (; Welsh language, Welsh for "dwarf dog") is one of two different varieties of livestock-herding dogs, herding dog breeds known as Welsh Corgi, Welsh corgis (originating in Wales), with the other being the Pembroke Welsh ...

Chesapeake Bay Retriever The Chesapeake Bay Retriever is a large dog breed, breed of dog belonging to the retriever, Gun dog, gundog, and sporting Breed Groups (dog), breed groups. The breed was developed in the United States Chesapeake Bay area during the 19th century. H ...

* pear- and deer-headed Chihuahuas * Chinese Crested * Chinook * Curly-Coated Retriever * Dalmatian *

Dhole The dhole ( ; ''Cuon alpinus'') is a canid native to South, East and Southeast Asia. It is anatomically distinguished from members of the genus ''Canis'' in several aspects: its skull is convex rather than concave in profile, it lacks a third ...

English Foxhound The English Foxhound is one of the four foxhound breeds of dog. It is a cousin of the American Foxhound. They are scent hounds, bred to hunt foxes by scent. Description Appearance The breed standards' guidelines for showing English Fo ...

* Field Spaniel * Finnish Lapphund *

Finnish Spitz Finnish may refer to: * Something or someone from, or related to Finland * Culture of Finland * Finnish people or Finns, the primary ethnic group in Finland * Finnish language, the national language of the Finnish people * Finnish cuisine See also ...

* Flat-Coated Retriever *

German Shorthaired Pointer The German Shorthaired Pointer or is a German breed of continental pointing dog of Braque type. It originated in the nineteenth century in what is now Germany. It is of medium size, and is an all-purpose gun dog suitable for hunting and retrie ...

German Wirehaired Pointer The German Wirehaired Pointer is a medium to large-sized Griffon (dog type), griffon type dog breed, breed of dog developed in the 19th century in Germany for versatile hunting. It became a leading gun dog in Germany in the later part of the 20th ...

* German Spitz * Golden Retriever * Irish Setter * Komondor *

Labrador Retriever The Labrador Retriever or simply Labrador or Lab is a British list of dog breeds, breed of water dog retriever gun dog. It was developed in the United Kingdom from St. John's water dogs imported from the Newfoundland Colony, colony of Newfoun ...

Miniature Pinscher The Miniature Pinscher, also known as the Zwergpinscher, Mini Pin and Min Pin for short, is a small breed of dog of the pinscher type originating in Germany. While the breed's visual similarities to the Toy Manchester Terrier, which somewhat resem ...

* Pomeranian *

Poodle The Poodle, called the in German () and the in French, is a breed of water dog. The breed is divided into four varieties based on size, the Standard Poodle, Medium Poodle, Miniature Poodle and Toy Poodle, although the Medium Poodle is no ...

(Miniature and Toy) * most

terrier Terrier () is a Dog type, type of dog originally bred to hunt vermin. A terrier is a dog of any one of many Dog breed, breeds or landraces of the terrier Dog type, type, which are typically small, wiry, Gameness, game, and fearless. There are fi ...

s * Mudi *

Pembroke Welsh Corgi The Pembroke Welsh Corgi (; Welsh for "dwarf dog") is a cattle herding dog breed that originated in Pembrokeshire, Wales. The name ''Corgi'' is of Welsh origin, and is a compound of the words and (mutated to ), meaning "dwarf" and "dog", r ...

* Puli * Rottweiler * Samoyed *

Siberian Husky The Siberian Husky is a dog breed, breed of medium-sized Working dog, working sled dog. The breed belongs to the Spitz genetic family. It is recognizable by its thickly furred Coat (dog), double coat, erect triangular ears, and distinctive mark ...

* St. Bernard * Vizsla *

Weimaraner The is a German breed of hunting dog of medium to large size, with history going back at least to the nineteenth century. It originated in the area of the city of Weimar (then in Saxe-Weimar-Eisenach, now in the state of Thuringia), for which ...

* Wirehaired Vizsla *

Xoloitzcuintle The Xoloitzcuintle (or Xoloitzquintle, Xoloitzcuintli, or Xolo) is one of several breeds of hairless dog. It is found in standard, intermediate, and miniature sizes. The Xolo also comes in a coated variety, totally covered in fur. Coated and hai ...

List of mesaticephalic cats

Note: Almost all domestic felines are mesaticephalic (“medium-headed”) * Abyssinian * American Shorthair * American Bobtail *

Bengal cat The Bengal cat is a breed of hybrid cat created from crossing of an Asian leopard cat (''Prionailurus bengalensis'') with domestic cats, especially the spotted Egyptian Mau. It is then usually bred with a breed that demonstrates a friendlier ...

Birman The Birman, also called the "Sacred Cat of Burma", is a domestic cat breed. The Birman is a long-haired, colour-pointed cat distinguished by a silky coat, deep blue eyes, and contrasting white "gloves" on each paw. The breed name is derive ...

* Bombay cat *

* Chartreux * Chausie * Colorpoint Shorthair *

Cymric cat The Cymric ( , ) is a Canadian cat breed. Some cat registries consider the Cymric a semi-long-haired variety of the Manx breed, rather than a separate breed. Except for the length of fur, in all other respects, the two varieties are the sam ...

Egyptian Mau The Egyptian Mau is a small to medium-sized, short to medium-haired List of cat breeds, cat breed. They are one of the few naturally spotted breeds of domesticated cat. The spots of the Mau occur on only the tips of the hairs of its coat. It is co ...

Felid hybrid A felid hybrid is any of a number of hybrids between various species of the cat family, Felidae. This article deals with hybrids between the species of the subfamily Felinae (feline hybrids). For hybrids between two species of the genus ''Pant ...

s * ''

Felis ''Felis'' is a genus of small and medium-sized cat species native to most of Africa and south of 60° latitude in Europe and Asia to Indochina. The genus includes the domestic cat. The smallest of the seven ''Felis'' species is the black-footed ...

'', or small cats * Maine Coon * Manx *

Munchkin cat The Munchkin is a breed of cat characterized by its very short legs, which are caused by a genetic mutation. Compared to many other cat breeds, it is a relatively new breed, documented since 1940s and officially recognized in 1991. The Munchkin ...

* Norwegian forest cat * Ocicat *

Pallas's cat The Pallas's cat (''Otocolobus manul''), also known as the manul, is a small wild cat with long and dense light grey fur, and rounded ears set low on the sides of the head. Its head-and-body length ranges from with a long bushy tail. It is w ...

Ragdoll The Ragdoll is a List of cat breeds, breed of cat with a distinct Point coloration, colorpoint coat and blue eyes. Its Morphology (biology), morphology is large and weighty, and it has a semi-long and silky soft coat. American breeder Ann Baker ...

Russian Blue The Russian Blue cat (), commonly referred to as just Russian Blue, is a pedigreed cat breed with solid blue colours that vary from a light shimmering silver to a darker, slate grey. The short, dense coat, which stands out from the body, has b ...

Russian White, Black and Tabby Russian(s) may refer to: *Russians (), an ethnic group of the East Slavic peoples, primarily living in Russia and neighboring countries *A citizen of Russia *Russian language, the most widely spoken of the Slavic languages *''The Russians'', a b ...

* Selkirk Rex * Siberian cat * Somali * Toyger *

Turkish Angora The Turkish Angora (, 'Ankara cat') is a breed of domestic cat. Turkish Angoras are one of the ancient, natural breeds of cat, having originated in central Anatolia ( Ankara Province in modern-day Turkey). The breed has been documented as ear ...

Turkish Van The Turkish Van is a semi-long-haired, standardised breed of domestic cat, which was developed in the United Kingdom from a selection of cats obtained from various cities of modern Turkey, especially southeast Turkey. The breed is rare, This s ...

List of mesaticephalic rabbits

* Dutch rabbit * Mini Rex * Polish rabbit *

New Zealand rabbit The New Zealand is a breed of rabbit, which despite the name, is American in origin. The breed originated in California, possibly from rabbits imported from New Zealand. New Zealand rabbits are available in five colors recognized by the Americ ...

American Sable The American Sable is a rabbit breed recognized by the American Rabbit Breeders Association (ARBA). The American Sable rabbit breed can trace its roots to colored throwbacks from purebred Chinchilla rabbits belonging to Otto Brock of San Gabriel ...

Other

* Aardwolf *

Alligator An alligator, or colloquially gator, is a large reptile in the genus ''Alligator'' of the Family (biology), family Alligatoridae in the Order (biology), order Crocodilia. The two Extant taxon, extant species are the American alligator (''A. mis ...

American black bear The American black bear (''Ursus americanus''), or simply black bear, is a species of medium-sized bear which is Endemism, endemic to North America. It is the continent's smallest and most widely distributed bear species. It is an omnivore, with ...

Brown bear The brown bear (''Ursus arctos'') is a large bear native to Eurasia and North America. Of the land carnivorans, it is rivaled in size only by its closest relative, the polar bear, which is much less variable in size and slightly bigger on av ...

Brown hyena The brown hyena (''Parahyaena brunnea''), also called the strandwolf, is a species of hyena found in Namibia, Botswana, western and southern Zimbabwe, southern Mozambique and South Africa. It is the only extant species in the genus ''Parahyaena' ...

Fur seal Fur seals are any of nine species of pinnipeds belonging to the subfamily Arctocephalinae in the family Otariidae. They are much more closely related to sea lions than Earless seal, true seals, and share with them external ears (Pinna (anatomy ...

Guinea pig The guinea pig or domestic guinea pig (''Cavia porcellus''), also known as the cavy or domestic cavy ( ), is a species of rodent belonging to the genus ''Cavia'', family Caviidae. Animal fancy, Breeders tend to use the name "cavy" for the ani ...

Leopard seal The leopard seal (''Hydrurga leptonyx''), also referred to as the sea leopard, is the second largest species of seal in the Antarctic (after the southern elephant seal). It is a top order predator, feeding on a wide range of prey including cep ...

Raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the North American, northern or common raccoon (also spelled racoon) to distinguish it from Procyonina, other species of raccoon, is a mammal native to North America. It is the largest ...

Sea lion Sea lions are pinnipeds characterized by external ear flaps, long foreflippers, the ability to walk on all fours, short and thick hair, and a big chest and belly. Together with the fur seals, they make up the family Otariidae, eared seals. ...

Sloth bear The sloth bear (''Melursus ursinus''), also known as the Indian bear, is a myrmecophagous bear species native to the Indian subcontinent. It feeds on fruits, ants and termites. It is listed as vulnerable on the IUCN Red List, mainly because of ...

Spotted hyena The spotted hyena (''Crocuta crocuta''), also known as the laughing hyena, is a hyena species, currently classed as the sole extant member of the genus ''Crocuta'', native to sub-Saharan Africa. It is listed as being of least concern by the IUC ...

Striped hyena The striped hyena (''Hyaena hyaena'') is a species of hyena native to North and East Africa, the Middle East, the Caucasus, Central Asia, and the Indian subcontinent. It is the only extant species in the genus ''Hyaena''. It is listed by the IU ...

Sun bear The sun bear (''Helarctos malayanus'') is a bear species in the Family (biology), family Ursidae found in the tropical forests of Southeast Asia. It is the only species in the genus ''Helarctos'' and the smallest bear species, standing nearly ...

Tasmanian devil The Tasmanian devil (''Sarcophilus harrisii''; palawa kani: ''purinina'') is a carnivorous marsupial of the family Dasyuridae. It was formerly present across mainland Australia, but became extinct there around 3,500 years ago; it is now con ...

Dolichocephalic animals

A dolichocephalic skull is relatively long-headed (typically with the breadth less than 80% or 75% of the length). Note: Almost all representatives of the infraphylum

Gnathostomata Gnathostomata (; from Ancient Greek: (') 'jaw' + (') 'mouth') are jawed vertebrates. Gnathostome diversity comprises roughly 60,000 species, which accounts for 99% of all extant vertebrates, including all living bony fishes (both ray-finned ...

(with rare exceptions) are dolichocephalic.

List of dolichocephalic canids

Note: Almost all canidae are dolichocephalic *

Afghan Hound The Afghan Hound is a hound distinguished by its thick, fine, silky coat, and a tail with a ring curl at the end. The breed originates in the cold mountains of Afghanistan. Its local name is () or (). Other names for this breed are Tāzī, Balkh ...

Airedale Terrier The Airedale Terrier (often shortened to "Airedale"), also called Bingley Terrier and Waterside Terrier, is a dog breed of the terrier type that originated in the valley (''dale'') of the River Aire, in the West Riding of Yorkshire, England. It ...

* Azawakh *

Basset Hound The Basset Hound is a short-legged breed of scent hound. The Basset Hound was developed in Great Britain from several now-extinct strains of France, French basset breeds. It was bred primarily for hunting rabbit and hare on foot, moving slowly en ...

Bedlington Terrier The Bedlington Terrier is a dog breed, breed of small dog named after the mining town of Bedlington, Northumberland in North East England. Originally bred to hunt, the Bedlington Terrier has since been used in dog racing, numerous List of dog s ...

Bloodhound The bloodhound is a large scent hound, originally bred for hunting deer, wild boar, rabbits, and since the Middle Ages, for tracking people. Believed to be descended from hounds once kept at the Abbey of Saint-Hubert, Belgium, in French it is ...

* Borzoi * Bull terrier * Cesky Terrier *

Coyote The coyote (''Canis latrans''), also known as the American jackal, prairie wolf, or brush wolf, is a species of canis, canine native to North America. It is smaller than its close relative, the Wolf, gray wolf, and slightly smaller than the c ...

Dachshund The dachshund ( or ; German: 'badger dog'), also known as the wiener dog or sausage dog, badger dog, doxen and doxie, is a short-legged, long-bodied, hound-type dog breed. The dog may be smooth-haired, wire-haired, or long-haired, with varie ...

* Doberman Pinscher *

Dingo The dingo (either included in the species ''Canis familiaris'', or considered one of the following independent taxa: ''Canis familiaris dingo'', ''Canis dingo'', or ''Canis lupus dingo'') is an ancient (basal (phylogenetics), basal) lineage ...

Fox Terrier Fox Terriers are two different breeds of the terrier dog type: the Smooth Fox Terrier and the Wire Fox Terrier. Both of these breeds originated in the 19th century from a handful of dogs who are descended from earlier varieties of United Kingdo ...

* Galgo Español *

German Shepherd Dog The German Shepherd, also known in Britain as an Alsatian, is a German breed of working dog of medium to large size. The breed was developed by Max von Stephanitz using various traditional German herding dogs from 1899. It was originally b ...

Great Dane The Great Dane is a German list of dog breeds, breed of large mastiff-sighthound, which descends from hunting dogs of the Middle Ages used to hunt bears, wild boar, and deer. They were also used as guardian dogs of German nobility. It is one o ...

Greyhound The English Greyhound, or simply the Greyhound, is a dog breed, breed of dog, a sighthound which has been bred for coursing, greyhound racing and hunting. Some are kept as show dogs or pets. Greyhounds are defined as a tall, muscular, smooth-c ...

* Irish Terrier * Irish Wolfhound * Italian Greyhound * Kangaroo hound * Kanni * Kerry Blue Terrier * Khalag Tazi * Long dog *

Lurcher A lurcher is a Dog crossbreed, crossbred dog resulting from dog reproduction, mating a greyhound or other sighthound with a dog of another Dog type, type such as a herding dog or a terrier. The lurcher is not a "breed," but is a generic descri ...

* Manchester Terrier * Miniature Bull Terrier * Peruvian Inca Orchid * Pharaoh Hound *

(Standard) * Rampur Greyhound * Rough Collie * Russian Black Terrier * Saluki * Schnauzer * Scottish Deerhound *

Scottish Terrier The Scottish Terrier (; also known as the Aberdeen Terrier), popularly called the Scottie, is a dog breed, breed of dog. Initially one of the highland breeds of terrier that were grouped under the name of ''Skye Terrier'', it is one of five br ...

* Sealyham Terrier * Serbian Hound * Shetland Sheepdog * Silken Windhound * Sloughi * Smooth Collie * Taigan * Welsh Terrier *

Whippet The Whippet is a British breed of dog of sighthound type. It closely resembles the Greyhound and the smaller Italian Greyhound, and is intermediate between them in size. In the nineteenth century it was sometimes called "the poor man's raceh ...

Wolf The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a Canis, canine native to Eurasia and North America. More than thirty subspecies of Canis lupus, subspecies of ''Canis lupus'' have been recognized, includin ...

List of dolichocephalic felines

* Balinese *

Devon Rex The Devon Rex is a tall-eared, short-haired breed of cat that emerged in England during the late 1950s. The breed is known for its atypical appearance, with an oddly shaped head, large eyes, and the short and wavy coat. History Origin Beryl Co ...

* Donskoy *

* Javanese *

Lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'', native to Sub-Saharan Africa and India. It has a muscular, broad-chested body (biology), body; a short, rounded head; round ears; and a dark, hairy tuft at the ...

Ocelot The ocelot (''Leopardus pardalis'') is a medium-sized spotted Felidae, wild cat that reaches at the shoulders and weighs between on average. It is native to the southwestern United States, Mexico, Central America, Central and South America, ...

* Oriental Bicolor and Tricolor *

Oriental Longhair The Oriental Longhair is a variety of domestic cat closely related to the Oriental Shorthair. The Oriental Longhair in some Cat registry, registries, such as The International Cat Association (TICA), it is considered a separate Cat breed, breed. ...

* Oriental Shorthair * ''Panthera'' hybrid * Peterbald * Sabertooth cats *

Savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...

* Siamese *

Snow leopard The snow leopard (''Panthera uncia'') is a species of large cat in the genus ''Panthera'' of the family Felidae. The species is native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because ...

* Sphynx *

Tiger The tiger (''Panthera tigris'') is a large Felidae, cat and a member of the genus ''Panthera'' native to Asia. It has a powerful, muscular body with a large head and paws, a long tail and orange fur with black, mostly vertical stripes. It is ...

List of dolichocephalic leporids

* English Spot * English Lop * Belgian Hare * All true

hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores and live Solitary animal, solitarily or in pairs. They nest in slight depressions called forms, and their young are precociality, able to fend for themselves ...

Other

Baboon Baboons are primates comprising the biology, genus ''Papio'', one of the 23 genera of Old World monkeys, in the family Cercopithecidae. There are six species of baboon: the hamadryas baboon, the Guinea baboon, the olive baboon, the yellow ba ...

* Bontebok *

Crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large, semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include ...

Cow Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are called co ...

Dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...

(including

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s) *

Dolphin A dolphin is an aquatic mammal in the cetacean clade Odontoceti (toothed whale). Dolphins belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontopori ...

* Domestic horse *

Donkey The donkey or ass is a domesticated equine. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a separate species, ''Equus asinus''. It was domes ...

Gharial The gharial (''Gavialis gangeticus''), also known as gavial or fish-eating crocodile, is a crocodilian in the family (biology), family Gavialidae and among the longest of all living crocodilians. Mature females are long, and males . Adult males ...

* Grevy's zebra *

Hartebeest The hartebeest (; ''Alcelaphus buselaphus''), also known as kongoni or kaama, is an Fauna of Africa, African antelope. It is the Monotypic taxon, only member of the genus ''Alcelaphus''. Eight subspecies have been described, including two som ...

Hyrax Hyraxes (), also called dassies, are small, stout, thickset, herbivorous mammals in the family Procaviidae within the order Hyracoidea. Hyraxes are well-furred, rotund animals with short tails. Modern hyraxes are typically between in length a ...

Kangaroo Kangaroos are marsupials from the family Macropodidae (macropods, meaning "large foot"). In common use, the term is used to describe the largest species from this family, the red kangaroo, as well as the antilopine kangaroo, eastern gre ...

Mule The mule is a domestic equine hybrid between a donkey, and a horse. It is the offspring of a male donkey (a jack) and a female horse (a mare). The horse and the donkey are different species, with different numbers of chromosomes; of the two ...

Onager The onager (, ) (''Equus hemionus''), also known as hemione or Asiatic wild ass, is a species of the family Equidae native to Asia. A member of the subgenus ''Asinus'', the onager was Scientific description, described and given its binomial name ...

Plains zebra The plains zebra (''Equus quagga'', formerly ''Equus burchellii'') is the most common and geographically widespread species of zebra. Its range is fragmented, but spans much of southern and eastern Africa south of the Sahara. Six or seven subspec ...

Polar bear The polar bear (''Ursus maritimus'') is a large bear native to the Arctic and nearby areas. It is closely related to the brown bear, and the two species can Hybrid (biology), interbreed. The polar bear is the largest extant species of bear ...

Thylacine The thylacine (; binomial name ''Thylacinus cynocephalus''), also commonly known as the Tasmanian tiger or Tasmanian wolf, was a carnivorous marsupial that was native to the Mainland Australia, Australian mainland and the islands of Tasmani ...

Warthog ''Phacochoerus'' is a genus in the family Suidae, commonly known as warthogs (pronounced ''wart-hog''). They are pigs who live in open and semi-open habitats, even in quite arid regions, in sub-Saharan Africa. The two species were formerly cons ...

Whale Whales are a widely distributed and diverse group of fully Aquatic animal, aquatic placental mammal, placental marine mammals. As an informal and Colloquialism, colloquial grouping, they correspond to large members of the infraorder Cetacea ...

Wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a Suidae, suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The speci ...

Wildebeest Wildebeest ( , ,), also called gnu ( or ), are antelopes of the genus ''Connochaetes'' and native to Eastern and Southern Africa. They belong to the family Bovidae, which includes true antelopes, cattle, goats, sheep, and other even-toed ...

References

External links

Cephalic index

Brachycephalic Experienced Veterinarians Database
{{DEFAULTSORT:Cephalic Index Biological anthropology