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''Meroktenos'' is a genus of basal
sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had l ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23  million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
that lived during the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
period of what is now
Lesotho Lesotho ( ), officially the Kingdom of Lesotho, is a country landlocked as an enclave in South Africa. It is situated in the Maloti Mountains and contains the highest mountains in Southern Africa. It has an area of over and has a populatio ...
.


Discovery and naming

In 1959,
François Ellenberger François () is a French masculine given name and surname, equivalent to the English name Francis. People with the given name * Francis I of France, King of France (), known as "the Father and Restorer of Letters" * Francis II of France, King ...
,
Paul Ellenberger Paul may refer to: *Paul (given name), a given name (includes a list of people with that name) * Paul (surname), a list of people People Christianity *Paul the Apostle (AD c.5–c.64/65), also known as Saul of Tarsus or Saint Paul, early Chri ...
, Jean Fabre and
Leonard Ginsburg Leonard or ''Leo'' is a common English masculine given name and a surname. The given name and surname originate from the Old High German ''Leonhard'' containing the prefix ''levon'' ("lion") from the Greek Λέων ("lion") through the Latin '' L ...
discovered the
type specimen In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally attached. In other words, a type is an example that serves to anchor or centralizes the ...
, a thighbone or femur and other assorted bones, south of the village of Thabana Morena. In 1962 these were addressed in a thesis by D. Costedoat. The exact location the bones were recovered, is today unknown. In 1993,
François-Xavier Gauffre François-Xavier is a French masculine given name. Notable people with the name include: * François-Xavier Archambault (1841–1893), a lawyer and political figure in Quebec * François-Xavier Audouin (1765–1837), a French clergyman and politi ...
assigned the remains to a second species of ''Melanorosaurus'': ''Melanorosaurus thabanensis''. The description was provisional, and in 1997 the fossil was described in more detail in a publication by
Jacques van Heerden Ancient and noble French family names, Jacques, Jacq, or James are believed to originate from the Middle Ages in the historic northwest Brittany region in France, and have since spread around the world over the centuries. To date, there are ov ...
and
Peter Malcolm Galton Peter Malcolm Galton (born 14 March 1942 in London) is a British vertebrate paleontologist who has to date written or co-written about 190 papers in scientific journals or chapters in paleontology textbooks, especially on ornithischian and prosau ...
. The specific name refers to the site Thabana-Morena in Lesotho. Gauffre assumed that the specimen had been found in the Upper Elliot Formation dating from the
Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic Period of the geologic timescale. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). The Hettangian follows the Rhaetian (part of the Triassi ...
-
Sinemurian In the geologic timescale, the Sinemurian is an age and stage in the Early or Lower Jurassic Epoch or Series. It spans the time between 199.3 ± 2 Ma and 190.8 ± 1.5 Ma (million years ago). The Sinemurian is preceded by the Hettangian an ...
and thus was about twenty million year younger than ''Melanorosaurus readi''. In 1996, he revised the date to the
Lower Elliot Formation The Elliot Formation is a geological formation and forms part of the Stormberg Group, the uppermost geological group that comprises the greater Karoo Supergroup. Outcrops of the Elliot Formation have been found in the northern Eastern Cape, sout ...
of the late Triassic in his non-published dissertation. He also referred the thighbone to a new genus and species ''
Thotobolosaurus ''Kholumolumo'' (referring to a type of dragon the local Basuto associate with dinosaurs), formerly "Kholumolumosaurus" or "Thotobolosaurus", is an extinct genus of massopodan sauropodomorph dinosaur, which was closely related to '' Sarahsauru ...
'' (now ''
Kholumolumo ''Kholumolumo'' (referring to a type of dragon the local Basuto associate with dinosaurs), formerly "Kholumolumosaurus" or "Thotobolosaurus", is an extinct genus of massopodan sauropodomorph dinosaur, which was closely related to '' Sarahsauru ...
''). This remained a non-valid ''nomen ex dissertatione'', as the name would never be published; furthermore the type material of this species does not coincide with that of ''M. thabanensis''.Gauffre F-X., 1996, ''Phylogénie des dinosaures prosauropodes et étude d’un prosauropode du Trias supérieur d’Afrique australe'' Dissertation, Muséum National d’Histoire Naturelle In 2016, ''M. thabanensis'' was appointed to the separate genus ''Meroktenos'' by
Claire Peyre de Fabrègues Clair or Claire may refer to: * Claire (given name), a list of people with the name Claire *Clair (surname) Places Canada * Clair, New Brunswick, a former village, now part of Haut-Madawaska * Clair Parish, New Brunswick * Pointe-Claire, ...
and Ronan Allain. The genus name is a combination of ancient Greek μηρός, ''meros'' ("thigh") and κτῆνος, ''ktènos'' ("beast"). The ''
combinatio nova ''Combinatio nova'', abbreviated ''comb. nov.'' (sometimes ''n. comb.''), is Latin for "new combination". It is used in taxonomic biology literature when a new name is introduced based on a pre-existing name. The term should not to be confused wi ...
'' thus becomes ''Meroktenos thabanensis'', the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
is the original ''Melanorosaurus thabanensis''. The
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
, MNHN.F.LES 16, consists of a right thighbone (MNHN.F.LES16c), a portion of the right ilium, with a piece of a vertebral
neural arch The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...
(MNHN.F.LES16a); a left pubic bone (MNHN.F.LES16b); and a second right metatarsal (MNHN.F.LES16d) associated with the skeleton. In 2016, a new specimen, MNHN.F.LES351, was referred to the species; consisting of a cervical vertebra, a left ulna and a, probably left, radius. It might have belonged to the same individual as the holotype, but this cannot be strictly proven.


Description

''Meroktenos'' has a femur length of around forty-eight centimeters, suggesting a body length of about four meters. In 2016, a revised list of distinguishing traits was given. The blade height of the ilium, measured from the highest point of the antitrochanter to the upper edge of the blade is 60% of the total height of the ilium, including peduncles. The rear blade of the ilium is roughly triangular in side view. The femur is very compact with a robusticity index, length divided by the circumference of the shaft, of 2.09. The femur has a straight shaft in both side and front views. The femoral shaft is substantially wider transversely than it is wide in side view, with a ratio of 1.58. On the rear of the femoral shaft, the fourth trochanter is oriented obliquely, running from the upper and inner side to the lower and outer side.


Phylogeny

In 2016, ''Meroktenos'' was placed in the
Sauropodomorpha Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had ...
, in a basal position. According to a
cladistic Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived ch ...
analysis, ''Meroktenos'' formed a polytomy with ''
Blikanasaurus ''Blikanasaurus'' is a genus of sauropodomorph dinosaur from the late Triassic of South Africa. The generic name ''Blikanasaurus'' is derived from Greek, meaning "lizard from Blikana". The species name ''cromptoni'' is taken from the surname of ...
'' and more derived taxa, above ''
Aardonyx ''Aardonyx'' (Afrikaans ''aard'', "earth" + Greek , "nail, claw") is a genus of basal sauropodomorph dinosaur. It is known from the type species ''Aardonyx celestae'' found from the Early Jurassic Elliot Formation of South Africa. ''A. celestae'' ...
'' in the evolutionary tree and below a polytomy including '' Camelotia'', ''
Melanorosaurus ''Melanorosaurus'' (meaning "Black Mountain Lizard", from the Greek ''melas/'', "black", ''oros/'', "mountain" + ''/'', "lizard") is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. A herbivore from South Afri ...
'' and more derived groups: A 2021 study by Pol and colleagues found ''Meroktenos'' to be a member of Lessemsauridae, being the sister taxon of a clade formed by ''
Kholumolumo ''Kholumolumo'' (referring to a type of dragon the local Basuto associate with dinosaurs), formerly "Kholumolumosaurus" or "Thotobolosaurus", is an extinct genus of massopodan sauropodomorph dinosaur, which was closely related to '' Sarahsauru ...
'' and ''
Ledumahadi ''Ledumahadi'' (meaning "giant thunderclap" in Sesotho language) is a genus of lessemsaurid sauropodomorph dinosaur from the Early Jurassic Elliot Formation in Free State Province, South Africa. The type and only species is ''L. mafube'', known ...
'':


Palaeobiology

The relative transverse width of the femur, the eccentricity, is remarkably high for such a small animal. These proportions were known previously only from
Sauropoda Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their ...
and explained as an adaptation to a very high absolute weight. Because the holotype probably was not a young animal and is unlikely to have attained giant proportions, the trait must have had a different function.


See also

*
2016 in paleontology Flora Plants Fungi Cnidarians Research * '' Yunnanoascus haikouensis'', previously thought to be a member of Ctenophora, is reinterpreted as a crown-group medusozoan by Han ''et al.'' (2016). * A study on the fossil corals from the Late T ...


Notes

{{Taxonbar, from=Q22665934 Sauropodomorphs Late Triassic dinosaurs of Africa Fossil taxa described in 2016 Paleontology in Lesotho