Menemerus Affinis
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''Menemerus affinis'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Menemerus ''Menemerus'' is a genus of Salticidae, jumping spiders that was first described by Eugène Simon, Eugène Louis Simon in 1868. They are long, flattened in shape, and very hairy, usually with brown and grayish hairs. Most species have white edges ...
'' that lives in the United Arab Emirates. The spider is externally hard to tell apart from the related '' Menemerus arabicus'' or '' Menemerus fagei''; this affinity is reflected in the species name. The spider was first described in 2010 by
Wanda Wesołowska Wanda Wesołowska (born 11 August 1950) is a Polish zoologist known for her work with jumping spiders. She has described more species of jumping spider than any contemporary writer, and is second only to Eugène Simon in the history of arachnolo ...
and Antonius van Harten based on a
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
discovered in
Ras Al Khaimah Ras Al Khaimah (; ), often referred to its initials RAK, is the largest city and capital of the Emirate of Ras Al Khaimah in the U.A.E. The city had a population of 191,753 in 2025, and is the sixth-most populous city in UAE after Dubai, Abu ...
. Other examples have been found in
Dubai Dubai (Help:IPA/English, /duːˈbaɪ/ Help:Pronunciation respelling key, ''doo-BYE''; Modern Standard Arabic, Modern Standard Arabic: ; Emirati Arabic, Emirati Arabic: , Romanization of Arabic, romanized: Help:IPA/English, /diˈbej/) is the Lis ...
. The spider is medium sized with a brown
cephalothorax The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cepha ...
that is between long and a greyish-brown
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
between long. It has brown
legs A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element cap ...
. The female is larger than the male and has an
epigyne The epigyne or epigynum is the external genital structure of female spiders. As the epigyne varies greatly in form in different species, even in closely related ones, it often provides the most distinctive characteristic for recognizing species. ...
with a large pocket to the front and a notch at the back. The male copulatory organs has an
embolus An embolus (; : emboli; from the Greek ἔμβολος "wedge", "plug") is an unattached mass that travels through the circulatory system, bloodstream and is capable of creating blockages. When an embolus Vascular occlusion, occludes a blood vess ...
paired to a large secondary conductor and two spikes, or apophyses, on the
pedipalp Pedipalps (commonly shortened to palps or palpi) are the secondary pair of forward appendages among Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to ...
tibia.


Taxonomy

''Menemerus affinis'' is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
jumping spider Jumping spiders are a group of spiders that constitute the family (biology), family Salticidae. , this family contained over 600 species description, described genus, genera and over 6,000 described species, making it the largest family of spide ...
that was first described by
Wanda Wesołowska Wanda Wesołowska (born 11 August 1950) is a Polish zoologist known for her work with jumping spiders. She has described more species of jumping spider than any contemporary writer, and is second only to Eugène Simon in the history of arachnolo ...
and Antonius van Harten in 2010. It is one of over 500
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
identified by the Polish
arachnologist Arachnology is the scientific study of arachnids, which comprise spiders and related invertebrates such as scorpions, pseudoscorpions, harvestmen, ticks, and mites. Those who study spiders and other arachnids are arachnologists. More narrowly ...
Wesolowska during her career. They allocated the spider to the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Menemerus ''Menemerus'' is a genus of Salticidae, jumping spiders that was first described by Eugène Simon, Eugène Louis Simon in 1868. They are long, flattened in shape, and very hairy, usually with brown and grayish hairs. Most species have white edges ...
'', first
circumscribed In geometry, a circumscribed circle for a set of points is a circle passing through each of them. Such a circle is said to ''circumscribe'' the points or a polygon formed from them; such a polygon is said to be ''inscribed'' in the circle. * Circum ...
in 1868 by
Eugène Simon Eugène Louis Simon (; 30 April 1848 – 17 November 1924) was a French naturalist who worked particularly on insects and spiders, but also on birds and plants. He is by far the most prolific spider Taxonomy (biology), taxonomist in history, ...
, that contains over 60 species. The genus name derives from two Greek words, meaning "certainly" and " diurnal".
Phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analysis has shown that the genus is related to the genera ''
Helvetia Helvetia () is a national personification of Switzerland, officially , the Swiss Confederation. The allegory is typically pictured in a flowing clothing, with a spear and a shield emblazoned with the Flag of Switzerland, Swiss flag, and commo ...
'' and ''
Phintella ''Phintella'' is a genus of Salticidae, jumping spiders that was first described by W. Bösenberg & Embrik Strand in 1906. Species it contains fifty-nine species and one subspecies, found in Oceania, Asia, Europe, and Africa: *''Phintella abnorm ...
''. The genus also shares some characteristics with the genera ''
Hypaeus ''Hypaeus'' is a genus of the spider family Salticidae (jumping spiders). Species , the World Spider Catalog accepted the following species: *'' Hypaeus annulifer'' Simon, 1900 – Brazil *'' Hypaeus barromachadoi'' Caporiacco, 1947 – Guyana ...
'' and ''
Pellenes ''Pellenes'' is a genus of jumping spiders that was first described by Eugène Louis Simon in 1876. It is considered a senior synonym of ''Hyllothyene''. They are dark to black with white stripes on the back, and often have bright red markings. ...
''. It is a member of the
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Heliophaninae Chrysillini is a tribe of jumping spider in the family Salticidae. In Maddison's 2015 revision of the family, the subfamily Heliophaninae was reclassified as a junior synonym of Chrysillini. Genera * '' Afraflacilla'' * ''Augustaea'' * ''Ch ...
, renamed
Chrysillini Chrysillini is a tribe of jumping spider in the family Salticidae. In Maddison's 2015 revision of the family, the subfamily Heliophaninae was reclassified as a junior synonym of Chrysillini. Genera * '' Afraflacilla'' * '' Augustaea'' * '' ...
by
Wayne Maddison Wayne Paul Maddison (born 1958) is a Canadian evolutionary biologist, arachnologist, and biological illustrator. He is Canada Research Chair in Biodiversity and a professor at the departments of zoology and botany at the University of British ...
in 2015. Chrysillines are
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
. The tribe is ubiquitous across most of the continents of the world. It is allocated to the subclade Saltafresia in the
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
Salticoida Salticoida is an unranked clade of the jumping spider family Salticidae. It is the larger and more widespread of the two subdivisions of the "typical" jumping spiders (subfamily Salticinae), occurring effectively world-wide. Its sister clade is ...
. In 2016,
Jerzy Prószyński Jerzy Prószyński (born 1935 in Warsaw) is a Polish arachnologist specializing in systematics of jumping spiders (family Salticidae). He is a graduate of the University of Warsaw, a long-term employee of the Siedlce University of Natural Sc ...
created a group of genera named Menemerines after the genus. The vast majority of the species in Menemerines are members of the genus, with additional examples from '' Kima'' and ''
Leptorchestes ''Leptorchestes'' is a genus (biology), genus of jumping spiders in the family Salticidae. As in several other genera of salticids, it ant mimicry, mimicks ants. Species * ''Leptorchestes algerinus'' Wanda Wesołowska, Wesołowska & Szeremeta, 2 ...
''. The species is named for its affinity with related species in the genus.


Description

''Menemerus affinis'' is a medium-sized spider. Its body is divided into two main parts, a
cephalothorax The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cepha ...
and an oval
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
. The male cephalothorax measures typically in length and in width with an abdomen typically long and wide. Males of this species have a
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
that is brown with a line down the middle formed of white hairs. The same hairs cover the darker eye field which is darker. The spider's face, known as the
clypeus The clypeus is one of the sclerites that make up the face of an arthropod. In insects, the clypeus delimits the lower margin of the face, with the labrum articulated along the ventral margin of the clypeus. The mandibles bracket the labrum, but ...
, brown, as are the mouthparts, consisting of
chelicerae The chelicerae () are the arthropod mouthparts, mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated ...
, labium and maxilae. The
sternum The sternum (: sternums or sterna) or breastbone is a long flat bone located in the central part of the chest. It connects to the ribs via cartilage and forms the front of the rib cage, thus helping to protect the heart, lungs, and major bl ...
, the underside of the cephalothorax, is also brown. The abdomen primarily greyish-brown and covered with dense long brown and greyish hairs with a wide leaf-like shape in the middle and a darker patch to the front. Its sides and underside are light. The spider's
spinneret A spinneret is a silk-spinning organ of a spider or the larva of an insect. Some adult insects also have spinnerets, such as those borne on the forelegs of Embioptera. Spinnerets are usually on the underside of a spider's opisthosoma, and ar ...
s are light grey. Its
legs A leg is a weight-bearing and locomotive anatomical structure, usually having a columnar shape. During locomotion, legs function as "extensible struts". The combination of movements at all joints can be modeled as a single, linear element cap ...
are brown, the foremost being darker than the rearmost pair, with dense brown leg hairs. The spider makes sounds using stridulatory apparatus of bristles on the legs and a grater on the chelicerae. A key feature of the male ''Menemerus affinis'' is its
embolus An embolus (; : emboli; from the Greek ἔμβολος "wedge", "plug") is an unattached mass that travels through the circulatory system, bloodstream and is capable of creating blockages. When an embolus Vascular occlusion, occludes a blood vess ...
, part of the
reproductive system The reproductive system of an organism, also known as the genital system, is the biological system made up of all the anatomical organs involved in sexual reproduction. Many non-living substances such as fluids, hormones, and pheromones are al ...
, which is short and slightly curved towards the
palpal bulb The two palpal bulbs – also known as palpal organs and genital bulbs – are the copulatory organs of a male spider. They are borne on the last segment of the pedipalps (the front "limbs" of a spider), giving the spider an appearance often descr ...
. The embolus is paired to a large secondary conductor that accompanies the embolus, which is
sclerotized Sclerosis (also sclerosus in the Latin names of a few disorders) is a hardening of tissue and other anatomical features. It may refer to: * Sclerosis (medicine), a hardening of tissue * in zoology, a process which forms sclerites, a hardened exo ...
at its base. Its
pedipalp Pedipalps (commonly shortened to palps or palpi) are the secondary pair of forward appendages among Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to ...
, which is brown with white hairs, has a tibia with two spikes, or apophyses. These run next to each other, facing upwards. The female is larger than the male, with a cephalothorax typically in length and in width and an abdomen typically and wide. The female is otherwise similar externally to the male, differing in only a few superficial aspects, such as there being wide white streaks on the side of the carapace. The copulatory organs are again characteristic of the species. The
epigyne The epigyne or epigynum is the external genital structure of female spiders. As the epigyne varies greatly in form in different species, even in closely related ones, it often provides the most distinctive characteristic for recognizing species. ...
is wide with a deep notch at the back and a large pocket to the front. There are two depressions to the sides. Internally, the structure is typical for the genus. The copulatory openings are hidden in large cups that have strong sclerotization. The insemination ducts are short and lead to spherical
spermatheca The spermatheca (pronounced : spermathecae ), also called ''receptaculum seminis'' (: ''receptacula seminis''), is an organ of the female reproductive tract in insects, e.g. ants, bees, some molluscs, Oligochaeta worms and certain other in ...
e. Spiders of the ''Menemerus'' genus are difficult to distinguish. Apart from its copulatory organs, ''Menemerus affinis'' is particularly hard to tell apart from the related species '' Menemerus arabicus'' or '' Menemerus fagei''. The similarity in their size and the design of their bodies makes them easy to confuse.


Distribution

''Menemerus'' spiders are found throughout Africa and Asia, and have been identified as far as Latin America. ''Menemerus affinis'' is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the United Arab Emirates. It was first found in the
Emirate of Ras Al Khaimah Ras Al Khaimah is the northernmost of the Emirates of the United Arab Emirates, seven emirates that make up the United Arab Emirates. The city of Ras Al Khaimah, abbreviated to RAK or RAK City, is the capital of the emirate and home to most of ...
, with the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
collected in 2008 in Wadi Shawka. Other examples are also found living in the
Emirate of Dubai The Emirate of Dubai is one of the seven emirates of the United Arab Emirates. It is the most populous emirate of the UAE. The capital of the emirate is the eponymous city, Dubai. Governance Dubai is governed as an absolute monarchy by the A ...
.


References


Citations


Bibliography

* * * * * * * * {{Taxonbar, from=Q3305803 Endemic fauna of the United Arab Emirates Salticidae Spiders described in 2010 Spiders of the Arabian Peninsula Taxa named by Wanda Wesołowska