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biology Biology is the scientific study of life and living organisms. It is a broad natural science that encompasses a wide range of fields and unifying principles that explain the structure, function, growth, History of life, origin, evolution, and ...
, mating is the pairing of either opposite-sex or hermaphroditic
organism An organism is any life, living thing that functions as an individual. Such a definition raises more problems than it solves, not least because the concept of an individual is also difficult. Many criteria, few of them widely accepted, have be ...
s for the purposes of
sexual reproduction Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote tha ...
. ''
Fertilization Fertilisation or fertilization (see American and British English spelling differences#-ise, -ize (-isation, -ization), spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give ...
'' is the fusion of two
gametes A gamete ( ) is a haploid cell that fuses with another haploid cell during fertilization in organisms that reproduce sexually. Gametes are an organism's reproductive cells, also referred to as sex cells. The name gamete was introduced by the Ge ...
. '' Copulation'' is the union of the
sex organ A sex organ, also known as a reproductive organ, is a part of an organism that is involved in sexual reproduction. Sex organs constitute the primary sex characteristics of an organism. Sex organs are responsible for producing and transporting ...
s of two sexually reproducing animals for
insemination Insemination is the introduction of sperm (in semen) into a female or hermaphrodite's reproductive system in order to fertilize the ovum through sexual reproduction. The sperm enters into the uterus of a mammal or the oviduct of an oviparous (eg ...
and subsequent
internal fertilization Internal fertilization is the union of an egg and sperm cell during sexual reproduction inside the female body. Internal fertilization, unlike its counterpart, external fertilization, brings more control to the female with reproduction. For inte ...
. Mating may also lead to external fertilization, as seen in
amphibians Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
, fishes and plants. For most species, mating is between two individuals of opposite sexes. However, for some hermaphroditic species, copulation is not required because the parent organism is capable of self-fertilization (
autogamy Autogamy or self-fertilization refers to the Cell fusion, fusion of two gametes that come from one individual. Autogamy is predominantly observed in the form of self-pollination, a Reproduction, reproductive mechanism employed by many flowering pl ...
); for example, banana slugs. The term ''mating'' is also applied to related processes in bacteria,
archaea Archaea ( ) is a Domain (biology), domain of organisms. Traditionally, Archaea only included its Prokaryote, prokaryotic members, but this has since been found to be paraphyletic, as eukaryotes are known to have evolved from archaea. Even thou ...
and viruses. Mating in these cases involves the pairing of individuals, accompanied by the pairing of their homologous chromosomes and then exchange of genomic information leading to formation of recombinant progeny (see mating systems).


Animals

For animals, mating strategies include
random mating Panmixia (or panmixis) means uniform random fertilization, which means individuals do not select a mate based on physical traits. A panmictic population is one where all potential parents may contribute equally to the gamete pool, and that these ga ...
, disassortative mating, assortative mating, or a
mating pool In biology, mating is the pairing of either opposite-sex or hermaphrodite, hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gamete, gametes. ''Copulation (zoology), Copulation'' is the un ...
. In some birds, it includes behaviors such as
nest A nest is a structure built for certain animals to hold Egg (biology), eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of ...
-building and feeding offspring. The human practice of mating and artificially inseminating domesticated animals is part of
animal husbandry Animal husbandry is the branch of agriculture concerned with animals that are raised for meat, animal fiber, fibre, milk, or other products. It includes day-to-day care, management, production, nutrition, selective breeding, and the raising ...
. In some terrestrial
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s, including insects representing basal (primitive)
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
clades, the male deposits
spermatozoa A spermatozoon (; also spelled spermatozoön; : spermatozoa; ) is a motile sperm cell (biology), cell produced by male animals relying on internal fertilization. A spermatozoon is a moving form of the ploidy, haploid cell (biology), cell that is ...
on the substrate, sometimes stored within a special structure.
Courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage or committed romantic, ''de facto'' relationship. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marri ...
involves inducing the female to take up the sperm package into her genital opening without actual copulation. Courtship is often facilitated through forming groups, called leks, in flies and many other insects. For example, male ''
Tokunagayusurika akamusi ''Tokunagayusurika akamusi'' is a midge fly species of the family ''Chironomidae'', commonly called "nonbiting midges" or "lake flies." Described by Tokunaga in 1938, the species is common to Trophic state index, eutrophic lakes in Japan. The m ...
'' forms swarms dancing in the air to attract females. In groups such as
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
and many spiders, males extrude sperm into secondary copulatory structures removed from their genital opening, which are then used to inseminate the female (in dragonflies, it is a set of modified sternites on the second abdominal segment; in spiders, it is the male
pedipalp Pedipalps (commonly shortened to palps or palpi) are the secondary pair of forward appendages among Chelicerata, chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to ...
s). In advanced groups of insects, the male uses its
aedeagus An aedeagus ( or aedeagi) is a reproductive organ of male arthropods through which they secrete sperm from the testes during copulation (zoology), copulation with a female. It can be thought of as the insect equivalent of a mammal's penis, th ...
, a structure formed from the terminal segments of the abdomen, to deposit sperm directly (though sometimes in a capsule called a "
spermatophore A spermatophore, from Ancient Greek σπέρμα (''spérma''), meaning "seed", and -φόρος (''-phóros''), meaning "bearing", or sperm ampulla is a capsule or mass containing spermatozoa created by males of various animal species, especiall ...
") into the female's reproductive tract. Other animals reproduce sexually with external fertilization, including many basal
vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...
s. Vertebrates reproduce with internal fertilization through
cloaca A cloaca ( ), : cloacae ( or ), or vent, is the rear orifice that serves as the only opening for the digestive (rectum), reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles, birds, cartilagin ...
l copulation (in reptiles, some fish, and most birds) or penile- vaginal penetration and
ejaculation Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the penis through the urethra. It is the final stage and natural objective of male sexual stimulation, and an essential component of natural conception. ...
of
semen Semen, also known as seminal fluid, is a bodily fluid that contains spermatozoon, spermatozoa which is secreted by the male gonads (sexual glands) and other sexual organs of male or hermaphrodite, hermaphroditic animals. In humans and placen ...
(in
mammals A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle e ...
). In domesticated animals, there are various type of mating methods being employed to mate animals like pen mating (when female is moved to the desired male into a pen) or paddock mating (where one male is let loose in the paddock with several females). File:Bagmati River, Pashupatinath, Nepal Animal sex バグマティ川とパシュパティナート火葬場 サルの交尾 5835.JPG, Macaque monkeys mating File:オオカミ(Gray wolf) (5339403526).jpg, Gray wolves mating File:LionsMating.jpg, Lions mating File:Snails mating.jpg, Hermaphroditic snails (''
Cornu aspersum ''Cornu aspersum'' (syn. ''Helix aspersa'', ''Cryptomphalus aspersus''), known by the common name garden snail, is a species of land snail in the family Helicidae, which includes some of the most familiar land snails. Of all terrestrial molluscs, ...
'')
mating In biology, mating is the pairing of either opposite-sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. '' Copulation'' is the union of the sex organs of two sexually repr ...
File:Tortoise mating.jpg, African spurred tortoises ('' Centrochelys sulcata'') mating File:Chalkhill blue butterflies (Polyommatus coridon) mating 1.jpg, Chalkhill blue butterflies ('' Lysandra coridon'') mating File:Hoverflies mating midair.jpg,
Hoverflies Hoverflies, also called flower flies or syrphids, make up the insect family (biology), family Syrphidae. As their common name suggests, they are often seen Hover (behaviour), hovering or nectaring at flowers; the adults of many species feed main ...
('' Simosyrphus grandicornis'') mating in midair File:Joined moths.JPG, Poplar hawk-moths ('' Laothoe populi'') mating File:Ladybird-Coccinellidae-mating.jpg,
Ladybug Coccinellidae () is a widespread family of small beetles. They are commonly known as ladybugs in North America and ladybirds in the United Kingdom; "lady" refers to mother Mary. Entomologists use the names ladybird beetles or lady beetles ...
s mating File:Aphrophora alni mating.jpg, Spittlebugs (''
Aphrophora alni ''Aphrophora alni'', the European alder spittle bug, is a species belonging to the family Aphrophoridae. Distribution This species is quite common and widespread. It is present in most of Europe, in the eastern Palearctic realm, in the Near East ...
'') mating File:Dogs mating.jpg,
Dogs The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers ...
mating File:Goats mating.jpg,
Goats The goat or domestic goat (''Capra hircus'') is a species of goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the famil ...
mating File:Two cats mating (September 2021).jpg,
Cats The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have shown that the ...
mating File:Gråsparv (Passer domesticus) - Ystad-2024.jpg, House sparrows mating. File:Sexually explicit flies..jpg, Fruit Flies mating


Plants and fungi

Like in animals, mating in other Eukaryotes, such as plants and
fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
, denotes . However, in vascular plants this is mostly achieved without physical contact between mating individuals (see
pollination Pollination is the transfer of pollen from an anther of a plant to the stigma (botany), stigma of a plant, later enabling fertilisation and the production of seeds. Pollinating agents can be animals such as insects, for example bees, beetles or bu ...
), and in some cases, e.g., in fungi no distinguishable male or female organs exist (see
isogamy Isogamy is a form of sexual reproduction that involves Gamete, gametes of the same Morphology (biology), morphology (indistinguishable in shape and size), and is found in most Unicellular organism, unicellular eukaryotes. Because both gametes lo ...
); however, mating types in some fungal species are somewhat analogous to
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
in animals, and determine whether or not two individual isolates can mate. ''
Yeasts Yeasts are eukaryotic, single-celled microorganisms classified as members of the fungus kingdom. The first yeast originated hundreds of millions of years ago, and at least 1,500 species are currently recognized. They are estimated to constitu ...
'' are
eukaryotic The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms ...
microorganism A microorganism, or microbe, is an organism of microscopic scale, microscopic size, which may exist in its unicellular organism, single-celled form or as a Colony (biology)#Microbial colonies, colony of cells. The possible existence of unseen ...
s classified in the kingdom
Fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
, with 1,500
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
currently described. In general, under high stress conditions like
nutrient A nutrient is a substance used by an organism to survive, grow and reproduce. The requirement for dietary nutrient intake applies to animals, plants, fungi and protists. Nutrients can be incorporated into cells for metabolic purposes or excret ...
starvation,
haploid Ploidy () is the number of complete sets of chromosomes in a cell (biology), cell, and hence the number of possible alleles for Autosome, autosomal and Pseudoautosomal region, pseudoautosomal genes. Here ''sets of chromosomes'' refers to the num ...
cells will die; under the same conditions, however,
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, ...
cells of ''
Saccharomyces cerevisiae ''Saccharomyces cerevisiae'' () (brewer's yeast or baker's yeast) is a species of yeast (single-celled fungal microorganisms). The species has been instrumental in winemaking, baking, and brewing since ancient times. It is believed to have be ...
'' can undergo sporulation, entering sexual reproduction (
meiosis Meiosis () is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, the sperm or egg cells. It involves two rounds of division that ultimately result in four cells, each with only one c ...
) and produce a variety of haploid
spore In biology, a spore is a unit of sexual reproduction, sexual (in fungi) or asexual reproduction that may be adapted for biological dispersal, dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores fo ...
s, which can go on to mate (conjugate) and reform the
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, ...
.


Protists

Protists are a large group of diverse
eukaryotic The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms ...
microorganism A microorganism, or microbe, is an organism of microscopic scale, microscopic size, which may exist in its unicellular organism, single-celled form or as a Colony (biology)#Microbial colonies, colony of cells. The possible existence of unseen ...
s, mainly
unicellular A unicellular organism, also known as a single-celled organism, is an organism that consists of a single cell, unlike a multicellular organism that consists of multiple cells. Organisms fall into two general categories: prokaryotic organisms and ...
animals and plants, that do not form tissues. The earliest eukaryotes were likely protists. Mating and sexual reproduction are widespread among extant eukaryotes including protists such as ''
Paramecium ''Paramecium'' ( , , plural "paramecia" only when used as a Common name, vernacular name) is a genus of eukaryotic, unicellular ciliates, widespread in freshwater, brackish, and Ocean, marine environments. Paramecia are often abundant in stagna ...
'' and ''
Chlamydomonas ''Chlamydomonas'' ( ) is a genus of green algae consisting of about 150 species of unicellular organism, unicellular flagellates, found in stagnant water and on damp soil, in freshwater, seawater, and even in snow as "snow algae". ''Chlamydom ...
''. In many eukaryotic species, mating is promoted by sex pheromones including the protist '' Blepharisma japonicum.'' Based on a phylogenetic analysis, Dacks and Roger proposed that facultative sex was present in the common ancestor of all eukaryotes. However, to many biologists it seemed unlikely until recently, that mating and sex could be a primordial and fundamental characteristic of eukaryotes. A principal reason for this view was that mating and sex appeared to be lacking in certain
pathogen In biology, a pathogen (, "suffering", "passion" and , "producer of"), in the oldest and broadest sense, is any organism or agent that can produce disease. A pathogen may also be referred to as an infectious agent, or simply a Germ theory of d ...
ic protists whose ancestors branched off early from the eukaryotic family tree. However, several of these protists are now known to be capable of, or to recently have had, the capability for
meiosis Meiosis () is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, the sperm or egg cells. It involves two rounds of division that ultimately result in four cells, each with only one c ...
and hence mating. To cite one example, the common intestinal parasite '' Giardia intestinalis'' was once considered to be a descendant of a protist lineage that predated the emergence of meiosis and sex. However, ''G. intestinalis'' was recently found to have a core set of genes that function in meiosis and that are widely present among sexual eukaryotes. These results suggested that ''G. intestinalis'' is capable of meiosis and thus mating and sexual reproduction. Furthermore, direct evidence for meiotic recombination, indicative of mating and sexual reproduction, was also found in ''G. intestinalis''. Other protists for which evidence of mating and sexual reproduction has recently been described are parasitic protozoa of the genus ''
Leishmania ''Leishmania'' () is a genus of parasitic protozoans, single-celled eukaryotic organisms of the trypanosomatid group that are responsible for the disease leishmaniasis. The parasites are transmitted by sandflies of the genus '' Phlebotomus'' ...
'', ''
Trichomonas vaginalis ''Trichomonas vaginalis'' is an Anaerobic organism, anaerobic, flagellated protozoan parasite and the causative agent of a Sexually transmitted infection, sexually transmitted disease called trichomoniasis. It is the most common pathogenic protoz ...
'', and acanthamoeba. Protists generally reproduce asexually under favorable environmental conditions, but tend to reproduce sexually under stressful conditions, such as starvation or heat shock.


See also

*
Heterosexuality Heterosexuality is romantic attraction, sexual attraction, or Human sexual activity, sexual behavior between people of the opposite sex or gender. As a sexual orientation, heterosexuality is "an enduring pattern of emotional, romantic, and/or ...
*
Animal husbandry Animal husbandry is the branch of agriculture concerned with animals that are raised for meat, animal fiber, fibre, milk, or other products. It includes day-to-day care, management, production, nutrition, selective breeding, and the raising ...
*
Breeding in the wild Breeding in the wild is the natural process of animal reproduction occurring in the natural habitat of a given species. This terminology is distinct from animal husbandry Animal husbandry is the branch of agriculture concerned with animals ...
* Breeding season * Evolution of sex * Lordosis behavior * Mate choice copying *
Mating system A mating system is a way in which a group is structured in relation to sexual behaviour. The precise meaning depends upon the context. With respect to animals, the term describes which males and females mating, mate under which circumstances. Reco ...
*
Reproduction Reproduction (or procreation or breeding) is the biological process by which new individual organisms – "offspring" – are produced from their "parent" or parents. There are two forms of reproduction: Asexual reproduction, asexual and Sexual ...
*
Sex determination system A sex-determination system is a biological system that determines the development of sexual characteristics in an organism. Most organisms that create their offspring using sexual reproduction have two common sexes, males and females, and in ...
*
Sexual conflict Sexual conflict or sexual antagonism occurs when the two sexes have conflicting optimal fitness (biology), fitness strategies concerning reproduction, particularly over the mode and frequency of mating, potentially leading to an evolutionary arms ...
*
Sexual intercourse Sexual intercourse (also coitus or copulation) is a sexual activity typically involving the insertion of the Erection, erect male Human penis, penis inside the female vagina and followed by Pelvic thrust, thrusting motions for sexual pleasure ...


References


External links


Introduction to Animal Reproduction

Advantages of Sexual Reproduction
{{Sex (biology) Animal developmental biology Reproduction in animals Sexology Sexuality Ethology Fertility