Marmots are large
ground squirrel
Ground squirrels are rodents of the squirrel family (Sciuridae) that generally live on the ground or in burrows, rather than in trees like the tree squirrels. The term is most often used for the medium-sized ground squirrels, as the larger ones ar ...
s in the
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Marmota'', with 15 species living in Asia, Europe, and North America. These
herbivore
A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
s are active during the summer, when they can often be found in groups, but are not seen during the winter, when they
hibernate underground. They are the heaviest members of the
squirrel
Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
family.
Description
Marmots are large rodents with characteristically short but robust legs, enlarged claws which are well adapted to digging, stout bodies, and large heads and incisors to quickly process a variety of vegetation. While most species are various forms of earthen-hued brown, marmots vary in fur coloration based roughly on their surroundings. Species in more open habitat are more likely to have a paler color, while those sometimes found in well-
forested
A forest is an ecosystem characterized by a dense community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological functio ...
regions tend to be darker.
Total length varies typically from about and body mass averages about in spring in the smaller species and in autumn, at times exceeding , in the larger species.
The largest and smallest species are not clearly known.
[ In North America, on the basis of mean linear dimensions and body masses through the year, the smallest species appears to be the Alaska marmot and the largest is the ]Olympic marmot
The Olympic marmot (''Marmota olympus'') is a rodent in the squirrel family, Sciuridae. It occurs only in the U.S. state of Washington, at the middle elevations of the Olympic Peninsula. The closest relatives of this species are the hoary mar ...
. Some species, such as the Himalayan marmot and Tarbagan marmot in Asia, appear to attain roughly similar body masses to the Olympic marmot, but are not known to reach as high a total length as the Olympic species. In the traditional definition of hibernation
Hibernation is a state of minimal activity and metabolic reduction entered by some animal species. Hibernation is a seasonal heterothermy characterized by low body-temperature, slow breathing and heart-rate, and low metabolic rate. It is mos ...
, the largest marmots are considered the largest "true hibernators" (since larger "hibernators" such as bear
Bears are carnivoran mammals of the family (biology), family Ursidae (). They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats ...
s do not have the same physiological characteristics as obligate hibernating animals such as assorted rodent
Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s, bat
Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
s and insectivores).
Biology
Some species live in mountainous areas, such as the Alps
The Alps () are some of the highest and most extensive mountain ranges in Europe, stretching approximately across eight Alpine countries (from west to east): Monaco, France, Switzerland, Italy, Liechtenstein, Germany, Austria and Slovenia.
...
, northern Apennines
The Apennines or Apennine Mountains ( ; or Ἀπέννινον ὄρος; or – a singular with plural meaning; )Latin ''Apenninus'' (Greek or ) has the form of an adjective, which would be segmented ''Apenn-inus'', often used with nouns s ...
, Carpathians
The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe and Southeast Europe. Roughly long, it is the third-longest European mountain range after the Urals at and the Scandinavian Mountains ...
, Tatras, and Pyrenees
The Pyrenees are a mountain range straddling the border of France and Spain. They extend nearly from their union with the Cantabrian Mountains to Cap de Creus on the Mediterranean coast, reaching a maximum elevation of at the peak of Aneto. ...
in Europe; northwestern Asia; the Rocky Mountains
The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
, Black Hills, the Cascade
Cascade, or Cascading may refer to:
Science and technology Science
* Air shower (physics), a cascade (particle shower) of subatomic particles and ionized nuclei
** Particle shower, a cascade of secondary particles produced as the result of a high ...
and Pacific Ranges
The Pacific Ranges are the southernmost subdivision of the Coast Mountains portion of the Pacific Cordillera. Located entirely within British Columbia, British Columbia, Canada, they run northwest from the lower stretches of the Fraser River to B ...
, and the Sierra Nevada
The Sierra Nevada ( ) is a mountain range in the Western United States, between the Central Valley of California and the Great Basin. The vast majority of the range lies in the state of California, although the Carson Range spur lies primari ...
in North America; and the Deosai Plateau in Pakistan
Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
and Ladakh
Ladakh () is a region administered by India as a union territory and constitutes an eastern portion of the larger Kashmir region that has been the subject of a Kashmir#Kashmir dispute, dispute between India and Pakistan since 1947 and India an ...
in India. Other species prefer rough grassland and can be found widely across North America and the Eurasian Steppe
The Eurasian Steppe, also called the Great Steppe or The Steppes, is the vast steppe ecoregion of Eurasia in the temperate grasslands, savannas and shrublands biome. It stretches through Manchuria, Mongolia, Xinjiang, Kazakhstan, Siberia, Europea ...
. The slightly smaller and more social prairie dog
Prairie dogs (genus ''Cynomys'') are herbivorous burrowing Marmotini, ground squirrels native to the grasslands of North America. There are five recognized species of prairie dog: black-tailed prairie dog, black-tailed, white-tailed prairie dog ...
is not classified in the genus ''Marmota'', but in the related genus ''Cynomys''.
Marmots typically live in burrow
file:Chipmunk-burrow (exits).jpg, An eastern chipmunk at the entrance of its burrow
A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of Animal lo ...
s (often within rockpiles, particularly in the case of the yellow-bellied marmot
The yellow-bellied marmot (''Marmota flaviventer''), also known as the rock chuck, is a large, stout-bodied ground squirrel in the marmot genus. It is one of fourteen species of marmots, and is native to mountainous and semi-arid regions of south ...
), and hibernate there through the winter. Most marmots are highly social and use loud whistles to communicate with one another, especially when alarmed.
Marmots mainly eat greens and many types of grasses, berries
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone fruit, stone or pit (fruit), pit although many wikt:pip#Etymology 2, pips or seeds may be p ...
, lichen
A lichen ( , ) is a hybrid colony (biology), colony of algae or cyanobacteria living symbiotically among hypha, filaments of multiple fungus species, along with yeasts and bacteria embedded in the cortex or "skin", in a mutualism (biology), m ...
s, moss
Mosses are small, non-vascular plant, non-vascular flowerless plants in the taxonomic phylum, division Bryophyta (, ) ''sensu stricto''. Bryophyta (''sensu lato'', Wilhelm Philippe Schimper, Schimp. 1879) may also refer to the parent group bryo ...
es, root
In vascular plants, the roots are the plant organ, organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often bel ...
s, and flower
Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s.
Subgenera and species
The following is a list of all ''Marmota'' species recognized by Thorington and Hoffman plus the recently defined ''M. kastschenkoi''.[ They divide marmots into two ]subgenera
In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus.
In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
.
Some extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
species of marmots are recognized from the fossil record, for example:
*†''Marmota arizonae'', Arizona, U.S.
*†''Marmota minor'', Nevada, U.S.
*†''Marmota vetus'', Nebraska, U.S.
History and etymology
Marmots have been known since antiquity. Research by the French ethnologist
Ethnology (from the , meaning 'nation') is an academic field and discipline that compares and analyzes the characteristics of different peoples and the relationships between them (compare cultural, social, or sociocultural anthropology).
Scien ...
Michel Peissel claimed the story of the "Gold-digging ant
The gold-digging ant is a mythical insect described in classical and medieval bestiaries. They were dog- or fox-sized ants that dug up gold in sandy areas. Some versions of the '' Physiologus'' said they came from Ethiopia, while Herodotus claim ...
" reported by the Ancient Greek
Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
historian Herodotus
Herodotus (; BC) was a Greek historian and geographer from the Greek city of Halicarnassus (now Bodrum, Turkey), under Persian control in the 5th century BC, and a later citizen of Thurii in modern Calabria, Italy. He wrote the '' Histori ...
, who lived in the fifth century BCE, was founded on the golden Himalayan marmot of the Deosai Plateau and the habit of local tribes such as the Brokpa to collect the gold
Gold is a chemical element; it has chemical symbol Au (from Latin ) and atomic number 79. In its pure form, it is a brightness, bright, slightly orange-yellow, dense, soft, malleable, and ductile metal. Chemically, gold is a transition metal ...
dust excavated from their burrows.
Some historians believe that Strabo
Strabo''Strabo'' (meaning "squinty", as in strabismus) was a term employed by the Romans for anyone whose eyes were distorted or deformed. The father of Pompey was called "Gnaeus Pompeius Strabo, Pompeius Strabo". A native of Sicily so clear-si ...
's (''léōn mýrmīx''), and Agatharchides
Agatharchides or Agatharchus ( or , ''Agatharchos'') of Cnidus was a Greek historian and geographer (flourished 2nd century BC).
Life
Agatharchides is believed to have been born at Cnidus, hence his appellation. As Stanley M. Burstein notes, the ...
's (''myrmīkoléōn''), most probably are the marmot.
An anatomically accurate image of a marmot was printed and distributed as early as 1605 by Jacopo Ligozzi
Jacopo Ligozzi (1547–1627) was an Italian painter, illustrator, designer, and miniaturist. His art can be categorized as late-Renaissance and Mannerism, Mannerist styles.
Biography
Born in Verona, he was the son of the artist Giovanni Erma ...
, who was noted for his images of flora and fauna.
The etymology of the term "marmot" is uncertain. It may have arisen from the Gallo-Romance Gallo-Romance can refer to:
* Old Gallo-Romance, the Romance language spoken from around 600 to 900 AD.
* Gallo-Romance languages, a branch of the Romance language family, which includes in the narrowest sense the ''langues d'oïl'' and Franco-Prov ...
prefix ''marm-'', meaning to mumble or murmur (an example of onomatopoeia
Onomatopoeia (or rarely echoism) is a type of word, or the process of creating a word, that phonetics, phonetically imitates, resembles, or suggests the sound that it describes. Common onomatopoeias in English include animal noises such as Oin ...
). Another possible origin is postclassical Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
, , meaning "mountain mouse".
Beginning in 2010, Alaska celebrates February 2 as " Marmot Day", a holiday intended to observe the prevalence of marmots in that state and take the place of Groundhog Day
Groundhog Day (, , , ; Lunenburg County, Nova Scotia, Lunenburg, Nova Scotia: Daks Day) is a tradition observed regionally in the United States and Canada on February 2 of every year. It derives from the Pennsylvania Dutch superstition that if ...
.
Relationship to the Black Death
Some historians and paleogeneticists have postulated that the ''Yersinia pestis
''Yersinia pestis'' (''Y. pestis''; formerly ''Pasteurella pestis'') is a Gram-negative bacteria, gram-negative, non-motile bacteria, non-motile, coccobacillus Bacteria, bacterium without Endospore, spores. It is related to pathogens ''Yer ...
'' variant that caused the Black Death
The Black Death was a bubonic plague pandemic that occurred in Europe from 1346 to 1353. It was one of the list of epidemics, most fatal pandemics in human history; as many as people perished, perhaps 50% of Europe's 14th century population. ...
pandemic that struck Eurasia in the 14th century
The 14th century lasted from 1 January 1301 (represented by the Roman numerals MCCCI) to 31 December 1400 (MCD). It is estimated that the century witnessed the death of more than 45 million lives from political and natural disasters in both Euro ...
originated from a variant for which marmots in China were the natural reservoir
In Infection, infectious disease ecology and epidemiology, a natural reservoir, also known as a disease reservoir or a reservoir of infection, is the population of organisms or the specific environment in which an infectious pathogen naturally li ...
species.[The American Historical Review]
"The Four Black Deaths"
December 17, 2020. Accessed March 27, 2010.
References
External links
The Marmot Burrow
{{Authority control
Articles containing video clips
Rodents of Asia
Rodents of Europe
Rodents of North America
Extant Miocene first appearances
Taxa named by Johann Friedrich Blumenbach