Marcus Graham (entomologist)
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Marcus William Robert deVere Graham (25 March 1915 – 27 March 1995) was an English
entomologist Entomology (from Ancient Greek ἔντομον (''éntomon''), meaning "insect", and -logy from λόγος (''lógos''), meaning "study") is the branch of zoology that focuses on insects. Those who study entomology are known as entomologists. In ...
who specialised in the
Chalcidoidea Chalcid wasps (, , for their metallic colour) are insects within the superfamily Chalcidoidea, part of the order Hymenoptera. The superfamily contains some 22,500 known species, and an estimated total diversity of more than 500,000 species, m ...
superfamily of the
Hymenoptera Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typi ...
. He was associated with both
Oxford University The University of Oxford is a collegiate research university in Oxford, England. There is evidence of teaching as early as 1096, making it the oldest university in the English-speaking world and the second-oldest continuously operating u ...
and
Trinity College, Dublin Trinity College Dublin (), officially titled The College of the Holy and Undivided Trinity of Queen Elizabeth near Dublin, and legally incorporated as Trinity College, the University of Dublin (TCD), is the sole constituent college of the Univ ...
.


Early life

Graham was born on 25 March 1915 in Rainton,
County Durham County Durham, officially simply Durham, is a ceremonial county in North East England.UK General Acts 1997 c. 23Lieutenancies Act 1997 Schedule 1(3). From legislation.gov.uk, retrieved 6 April 2022. The county borders Northumberland and Tyne an ...
, the son of the Reverend Robert Graham who was originally from
County Donegal County Donegal ( ; ) is a Counties of Ireland, county of the Republic of Ireland. It is in the Provinces of Ireland, province of Ulster and is the northernmost county of Ireland. The county mostly borders Northern Ireland, sharing only a small b ...
and Ms Bertha Heaton of
Windermere Windermere (historically Winder Mere) is a ribbon lake in Cumbria, England, and part of the Lake District. It is the largest lake in England by length, area, and volume, but considerably smaller than the List of lakes and lochs of the United Ki ...
. He developed an interest in entomology as a boy. He enlisted in the British Army at the start of
World War II World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...
and served in India from 1942 until the end of the war.


Entomology

Graham began to publish entomological papers in 1941. He entered Trinity College to study for a degree in Natural Science at the end of 1945, graduating in 1950 with both a
Bachelor of Arts A Bachelor of Arts (abbreviated B.A., BA, A.B. or AB; from the Latin ', ', or ') is the holder of a bachelor's degree awarded for an undergraduate program in the liberal arts, or, in some cases, other disciplines. A Bachelor of Arts deg ...
and a
Bachelor of Science A Bachelor of Science (BS, BSc, B.S., B.Sc., SB, or ScB; from the Latin ') is a bachelor's degree that is awarded for programs that generally last three to five years. The first university to admit a student to the degree of Bachelor of Scienc ...
degree, being awarded First Class Honours and a Gold Medal. While he was still a student he made the first recorded observation of the
sawfly Sawflies are wasp-like insects that are in the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plant ...
'' Harpiphorus lepidus'' in
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
. This species remains rare in Ireland. Also while in Ireland he was mentored by Arthur Wilson Stelfox and studied the
Braconidae The Braconidae are a family of parasitoid wasps. After the closely related Ichneumonidae, braconids make up the second-largest family in the order Hymenoptera, with about 17,000 recognized species and many thousands more undescribed. One analysis ...
, especially their taxonomy. He and Stelfox published several joint papers during this time which described several species new to science. However, Stelfox advised him to concentrate on the Chalcidoidea. Soon after returning to England he was appointed curator of the University Museum, Hope Department at Oxford University. In 1953 he married Eleanor Mary Scheppens, nicknamed Nora, and their only child, a son called John, was born in 1957. In 1955 he was awarded a D. Phil. degree by the University of Oxford, where as well as curating he was an administrator, teacher and researcher. He was a major contributor to the knowledge of the European Chalcid fauna, especially the
Pteromalidae The Pteromalidae are a large family of wasps, the majority being parasitoids of other insects. They are found throughout the world in virtually all habitats, and many are important as biological control agents. The oldest known fossil is known fr ...
,
Mymaridae The Mymaridae, commonly known as fairyflies or fairy wasps, are a family (biology), family of chalcid wasp, chalcidoid wasps found in temperate, subtropical, and tropical regions throughout the world. The family contains around 100 genera with 1, ...
and the
Eulophidae The Eulophidae is a large family of hymenopteran insects, with over 4,300 described species in some 300 genera. The family includes the genus '' Elasmus'', which used to be treated as a separate family, "Elasmidae", and is now treated as a subfa ...
. Nora was also interested in entomology and often joined Marcus on collecting expeditions and she collected the type specimen of the striking Braconid '' Chorebus norae'' on one such trip to
Madeira Madeira ( ; ), officially the Autonomous Region of Madeira (), is an autonomous Regions of Portugal, autonomous region of Portugal. It is an archipelago situated in the North Atlantic Ocean, in the region of Macaronesia, just under north of ...
, which Graham named in her honour. He retained his links with Dublin where he worked on Trinity College's Haliday Collection, carefully curating the collection to restore its scientific utility. The research he conducted during his visits to Trinity College enabled him to make many important discoveries and as a result of the field work he conducted in Ireland, he was able to add many species of
chalcid wasp Chalcid wasps (, , for their metallic colour) are insects within the superfamily Chalcidoidea, part of the order Hymenoptera. The superfamily contains some 22,500 known species, and an estimated total diversity of more than 500,000 species, m ...
to the list of those found in Ireland, some being new to science. Among the pteromalids he found in Ireland many have yet to be found in Britain. He retired from the Hope in 1981, and made his last visit to Dublin in 1984. During his career he described 60 new genera and 475 species of Hymenoptera.


Other interests

Outside of entomology Graham had a number of interests including languages, especially
Romance languages The Romance languages, also known as the Latin or Neo-Latin languages, are the languages that are Language family, directly descended from Vulgar Latin. They are the only extant subgroup of the Italic languages, Italic branch of the Indo-E ...
, mediaeval literature, history, naval architecture and painting. However, he continued to work on entomology after retirement and was preparing a paper on the revision of the genus ''
Torymus ''Torymus'' is a genus of chalcid wasps from the family Torymidae, and there are more than 400 spp. worldwide. It was named by the Swedish naturalist Johan Wilhelm Dalman in 1820. Most species are ectoparasitoids of gall forming insects, usuall ...
'' when he died, he refereed a paper for the ''Bulletin of the Irish Biogeographical Society'' just before he died and gave the authors invaluable advice, despite his failing health and hospitalisation. He died on 27 March 1995 and was survived by his wife, Nora, and son, John.


References

{{DEFAULTSORT:Graham, Marcus (entomologist) English entomologists Hymenopterists 1915 births 1995 deaths 20th-century English zoologists