Mansourasaurus
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''Mansourasaurus'' ("
Mansoura Mansoura (; ', , rural: ) is a city in Egypt located on the eastern bank of the Damietta branch of the Nile river. The city is the capital of the Dakahlia Governorate and has a population of 621,953 as of 2021. Etymology ''Mansoura'' in Arabic ...
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
herbivorous A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
lithostrotia Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch ''et al.'' in 2004 as the most recent common ancestor of ''Malawisaurus'' and ''Saltasaurus'' an ...
n
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the
Quseir Formation The Quseir Formation is a geological formation in the vicinity of the Kharga Oasis in Egypt.
of
Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...
. The
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * ...
and only species is ''Mansourasaurus shahinae''. The discovery of ''Mansourasaurus'' was considered quite significant by
paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
s, because very few
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
sauropod remains have been found in Africa, where the rocky
strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...
that preserve remains elsewhere and produce rich fossil beds were typically not found exposed at or near ground level.


Discovery and naming

Hesham Sallam, a paleontologist at
Mansoura University Mansoura University was founded in 1972 in Mansoura city, Egypt. It is in the middle of the Nile Delta. It is one of the biggest Egyptian universities and has contributed much to the cultural and scientific life in Mansoura and Egypt. History ...
, together with a team of students, discovered a sauropod skeleton in the
Dakhla Oasis Dakhla Oasis or Dakhleh Oasis ( Egyptian Arabic: , , "''the inner oases"''), is one of the seven oases of Egypt's Western Desert. Dakhla Oasis lies in the New Valley Governorate, 350 km (220 mi.) from the Nile and between the oases ...
in Egypt's Western Desert. In 2016, it was reported that over thirty dinosaur specimens had been excavated, among them titanosaurian sauropods.Iman El-Dawoudi, Patrick M O'Connor, Mahmoud Kora and Joseph J W Sertich, 2016, "NEW DINOSAUR REMAINS FROM THE CAMPANIAN QUSEIR FORMATION, WESTERN DESERT, EGYPT", ''SVP October 2016, Program and Abstracts'', p 129 Based on this skeleton, the type species ''Mansourasaurus shahinae'' was named and described in January 2018 by Hesham M. Sallam, Eric Gorscak, Patrick M. O'Connor, Iman A. El-Dawoudi, Sanaa El-Sayed, Sara Saber, Mahmoud A. Kora, Joseph J. W. Sertich, Erik R. Seiffert, and Matthew C. Lamanna. The generic name refers to the Mansoura University. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honours Mona Shahin, one of the founders of the Mansoura University Vertebrate Paleontology Center. The ''Mansourasaurus''
specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...
described in 2018 is its
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
, MUVP 200, discovered in a layer of the Quseir Formation dating from the late
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
, about seventy-three million years old. It consists of a partial skeleton with skull and lower jaws. It contains a fragment of the skull roof, a part of the lower
braincase In human anatomy, the neurocranium, also known as the braincase, brainpan, brain-pan, or brainbox, is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calv ...
, the dentaries of the lower jaws, three neck vertebrae, two back vertebrae, eight ribs, the right scapula, the right
coracoid A coracoid is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is present as part of the scapula, but this is n ...
, both humeri, a radius, a third
metacarpal In human anatomy, the metacarpal bones or metacarpus, also known as the "palm bones", are the appendicular bones that form the intermediate part of the hand between the phalanges (fingers) and the carpal bones ( wrist bones), which articulate ...
, three
metatarsal The metatarsal bones or metatarsus (: metatarsi) are a group of five long bones in the midfoot, located between the tarsal bones (which form the heel and the ankle) and the phalanges ( toes). Lacking individual names, the metatarsal bones are ...
s, and parts of
osteoderm Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amph ...
s. The skeleton was found on a surface of four by three metres. It was not articulated. The authors concluded that the holotype is a juvenile specimen because the bones of its shoulder girdle had not yet fused. An
ulna The ulna or ulnar bone (: ulnae or ulnas) is a long bone in the forearm stretching from the elbow to the wrist. It is on the same side of the forearm as the little finger, running parallel to the Radius (bone), radius, the forearm's other long ...
, specimen MUVP 201, found at twenty metres distance from the skeleton, was not referred to the species as it seemed somewhat too large for the holotype individual and a general connection to the species could not be proven.


Description

The not fully-grown holotype individual was about long. It probably weighed about 5,000 kg, approximately the same as a bull African elephant. The describing authors indicated a number of distinguishing traits. These are
autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to ...
, unique derived characters. Each lower jaw dentary bears ten teeth. Where the dentaries touch each other, at the front of the lower jaws, they possess a common "chin", equalling a third of the front height. The horizontal groove in the inner side of the dentary, the ''fossa Meckeliana'', largely opens to below. The anterior middle neck vertebrae are pierced by a
foramen In anatomy and osteology, a foramen (; : foramina, or foramens ; ) is an opening or enclosed gap within the dense connective tissue (bones and deep fasciae) of extant and extinct amniote animals, typically to allow passage of nerves, artery, ...
in the rear side. In at least one anterior middle neck vertebra, the parapophysis, the process which bears the facet for the lower rib head, has a horizontal length equal to the vertebral centrum as a whole. With some anterior neck vertebrae, the bone web between the heads of the neck rib is pierced by a foramen. The lower end of the radius has a transverse width four times larger than the width measured from the front to the rear.


Phylogeny

''Mansourasaurus'' was placed in the
Titanosauria Titanosaurs (or titanosaurians; members of the group Titanosauria) were a diverse group of Sauropoda, sauropod dinosaurs, including genera from all seven continents. The titanosaurs were the last surviving group of long-necked sauropods, with tax ...
in a derived position as a
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
of '' Lohuecotitan''. A
cladistic Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is ...
analysis showed it to belong to a
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
of otherwise largely Eurasian sauropods, also including ''
Ampelosaurus ''Ampelosaurus'' ( ; meaning "vine lizard") is a titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now France. Its type species is ''A. atacis'', named by Le Loeuff in 1995. Its remains were found in a level dating from ...
'', '' Lirainosaurus'', ''
Nemegtosaurus ''Nemegtosaurus'' (meaning 'Reptile from the Nemegt') was a sauropod dinosaur from the Late Cretaceous Period of what is now Mongolia. Measuring an estimated long and weighing , it was named after the Nemegt Basin in the Gobi Desert, where the ...
'', ''
Opisthocoelicaudia ''Opisthocoelicaudia'' is a genus of sauropod dinosaur of the Late Cretaceous Period discovered in the Gobi Desert of Mongolia. The type species is ''Opisthocoelicaudia skarzynskii''. A well-preserved skeleton lacking only the head and neck wa ...
'', and '' Paludititan'', more or less contemporaneous forms. Hypotheses about relationships between
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
African and Eurasian sauropods had been hard to test because very few of their remains had been found in Africa. ''Mansourasaurus'' represents the best-known continental African (i.e., excluding
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
) titanosaur of the
Upper Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cret ...
from the time period after the
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
. Its existence would show that the continent was far less isolated from the various Eurasian landmasses than had been assumed. The ancestors of ''Mansourasaurus'' would have reached Africa from Europe.


See also

*
2018 in paleontology Flora Plants Fungi Cnidarians Research * New three dimensionally Phosphatic fossilization, phosphatized Micropaleontology#Microfossils, microfossils of Crown jellyfish, coronate scyphozoan ''Qinscyphus necopinus'', including a new type of ...


References

{{Taxonbar, from=Q47518933 Lithostrotia Dinosaur genera Campanian dinosaurs Fossil taxa described in 2018 Dinosaurs of Egypt