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The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s. They are
cosmopolitan Cosmopolitan may refer to: Internationalism * World citizen, one who eschews traditional geopolitical divisions derived from national citizenship * Cosmopolitanism, the idea that all of humanity belongs to a single moral community * Cosmopolitan ...
in distribution. One of the first described species has a color pattern reminiscent of the red coats of early British soldiers, hence the common name. They are also known commonly as leatherwings because of their soft
elytra An elytron (; ; : elytra, ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometime ...
. Historically, these beetles were placed in a superfamily "Cantharoidea", which has been subsumed by the superfamily
Elateroidea The Elateroidea are a large superfamily of beetles. It contains the familiar click beetles, fireflies, and soldier beetles and their relatives. It consists of about 25,000 species. Description Elateroidea is a morphologically diverse group ...
; the name is still sometimes used as a rankless grouping, including the families Cantharidae,
Lampyridae The Lampyridae are a family of elateroid beetles with more than 2,000 described species, many of which are light-emitting. They are soft-bodied beetles commonly called fireflies, lightning bugs, or glowworms for their conspicuous production ...
,
Lycidae The Lycidae are a family in the beetle order Coleoptera, members of which are commonly called net-winged beetles. These beetles are cosmopolitan, being found in Nearctic, Palearctic, Neotropical, Afrotropical, Oriental, and Australian ecoregions ...
,
Omethidae Omethidae is a family of Elateroidea sometimes known as the false soldier beetles. They are native to South, Southeast and Eastern Asia and the Americas. Their biology is obscure and their larvae are unknown. They appear to inhabit vegetation in ...
(which includes Telegeusidae),
Phengodidae The beetle family Phengodidae is known also as glowworm beetles, whose larvae are known as glowworms. The females and larvae have bioluminescent organs. They occur throughout the New World from extreme southern Canada to Chile, numbering over ...
, and
Rhagophthalmidae The Rhagophthalmidae are a family of beetles within the superfamily Elateroidea. Members of this beetle family have bioluminescent organs on the larvae, and sometimes adults, and are closely related to the Phengodidae ( American glowworm beetles ...
. Soldier beetles often feed on nectar and pollen as well as predating other small insects. The larvae are caterpillar like, dark colored, active and covered in fine hairs, earning them the name velvet worms. They feed on the ground and in foliage hunting eggs, small insects, snails and other small creatures.


Evolutionary history

The oldest described member of the family is '' Molliberus'' from the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
(early
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) aged El Soplao amber from Cantabria, Spain, belonging to the tribe
Cantharini Cantharini is a tribe of soldier beetles in the family Cantharidae. There are more than 40 genera and over 500 described species in Cantharini. Genera These 46 genera belong to the tribe Cantharini: * ''Ancistronycha'' Märkel, 1852 * ''Armid ...
in the subfamily
Cantharinae Cantharinae is a subfamily of beetles in the family Cantharidae. There are at least 200 described species in Cantharinae. ;Taxonomic note: *Lawrence and Newton (1995) give the authorship of this subfamily as Imhoff, 1856 (1815). Tribes and gene ...
. Other described genera include 6 from the early
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(early
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) aged
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, with 5 belonging to Cantharinae and one to
Malthininae The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are Cosmopolitan distribution, cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the Red coat (Bri ...
, and '' Katyacantharis'', from the Cenomanian aged Agdzhakend amber of Azerbaijan, suggested to belong to Cantharinae. Indeterminate specimens have been reported from the
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
aged Koonwarra fossil bed of the
Strzelecki Group The Halibut Field is an oil field, within the Gippsland Basin offshore of the Australian state of Victoria (Australia), Victoria. The oil field is located approximately 64 km offshore of southeastern Australia. The total area of this field ...
, Australia and the
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
aged
Lebanese amber Lebanese amber is fossilized resin found in Lebanon and its surroundings. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropic ...
.


Subfamilies, tribes and selected genera

Five subfamilies are normally accepted:


Cantharinae Cantharinae is a subfamily of beetles in the family Cantharidae. There are at least 200 described species in Cantharinae. ;Taxonomic note: *Lawrence and Newton (1995) give the authorship of this subfamily as Imhoff, 1856 (1815). Tribes and gene ...

*
tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Cantharini ** ''
Cantharis ''Cantharis'' is a large genus of soldier beetles in the family Cantharidae with narrow and soft elytra. The poisonous Spanish fly is superficially similar and is associated with the scientific name ''Cantharis vesicatoria''. It is also sometimes ...
'' ** ''
Rhagonycha ''Rhagonycha''von Eschscholtz JF (1830) Nova genera Coleopterorum Faunae Europaeae. ''Bulletin de la Société Impériale des Naturalistes de Moscou'' 2(1): 63–66. is a genus of soldier beetle belonging to the family Cantharidae. There are at ...
'' * tribe Podabrini ** ''
Podabrus ''Podabrus'' is a genus of soldier beetles in the family Cantharidae. There are at least 110 described species in ''Podabrus'', recorded from Europe, North America and Asia. Many ''Podabrus'' appear to Batesian mimicry, mimic fireflies, such as ...
''


Chauliognathinae

* tribe Chauliognathini ** '' Belotus'' ** '' Chauliognathus'' * tribe Ichthyurini ** '' Trypherus''


Dysmorphocerinae

* '' Afronycha'' Wittmer, 1949 - central-southern Africa * '' Asilis'' Broun, 1893 - New Zealand * '' Compsonycha'' * '' Dysmorphocerus'' Solier, 1849 * '' Flabelloontelus'' * '' Geigyella'' Wittmer, 1972 - New Guinea * '' Hansasilis'' * '' Heteromastix'' Boheman, 1858 - Australia * '' Hyponotum'' * '' Micronotum'' * '' Neoontelus'' Wittmer, 1972 - New Zealand * '' Oontelus'' Solier, 1849 - S. America * '' Plectocephalon'' * ''
Plectonotum ''Plectonotum'' is a genus of soldier beetles in the family Cantharidae. There is at least one described species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating ...
'' Gorham, 1891 -
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...


Malthininae

* tribe Malchinini ** '' Macrocerus'' Motschulsky, 1845 - Europe (synonym ''Malchinus'') * tribe Malthinini ** '' Caccodes'' Sharp, 1885 - Central America, Pacific islands ** '' Malthinellus'' Kiesenwetter, 1874 - Japan ** '' Malthinus'' Latreille, 1805 - Japan, Europe, N. America * tribe Malthodini ** '' Frostia'' Bert. ex Guill. ** ''
Malthodes ''Malthodes'' is a genus of soldier beetles in the family Cantharidae The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are Cosmopolitan distribution, cosmopolitan in distribution. One of the fi ...
'' Kiesenwetter, 1852 - mostly Europe, N. America & New Zealand ** †'' Archaeomalthodes'' Hsiao et al. 2016 Burmese amber, Myanmar, Cenomanian


Silinae

* tribe Silini ** ''
Cordylocera ''Cordylocera'' is a genus of soldier beetles (insects in the family Cantharidae). Species * ''Cordylocera albovittata'' (Wittmer, 1964) * ''Cordylocera antennata'' Guérin-Méneville, 1830 * ''Cordylocera atricornis'' Guérin-Méneville, 183 ...
'' Guérin-Méneville, 1830 ** '' Silis'' Charpentier, 1825 * tribe Tytthonyxini ** ''
Tytthonyx ''Tytthonyx'' is a genus of soldier beetles in the family Cantharidae. There are about 19 described species in ''Tytthonyx''. Species These 19 species belong to the genus ''Tytthonyx'': * '' Tytthonyx bicolor'' LeConte, 1885 * '' Tytthonyx cuba ...
'' LeConte, 1851


Reproduction

Large males of the soldier beetle exercise choice for larger females. Body size correlates with the abilities of males to secure females, and of females to evade males.


See also

* List of Cantharidae genera


References


External links


Delta key

Keys for the identification of British Cantharidae
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