''Malayopython'' is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of constricting snakes in the
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Pythonidae
The Pythonidae, commonly known as pythons, are a family of nonvenomous snakes found in Africa, Asia, and Australia. Among its members are some of the largest snakes in the world. Ten genera and 42 species are currently recognized.
Distributi ...
. The genus is native to
India
India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
and
Southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
. It contains two
species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
, both of which were previously classified within the genus ''
Python''. However, multiple studies recovered these species as distinct. Known as the "''reticulatus'' clade", it was eventually found to be a
sister lineage
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree.
Definition
The expression is most easily illustrated by a cladogram:
Taxon A a ...
to a lineage giving rise to the Indo-Australian pythons rather than the genus ''Python''.
Taxonomy
In 1975, American herpetologist Samuel Booker McDowell divided the genus ''Python'' into a "''molurus'' group" and "''reticulatus'' group" on the basis of differences in
supralabial
In reptiles, the supralabial scales, also called upper-labials, are those scales that border the mouth opening along the upper jaw. They do not include the median scaleWright AH, Wright AA. 1957. Handbook of Snakes. Comstock Publishing Associates ...
pits (shallow diagonal slits in the latter, square or triangular in the former) and infralabial pits (shallow and not in a groove in the former, in a groove in the latter), as well as differences in the ectopterygoid and
hemipenis
A hemipenis (plural hemipenes) is one of a pair of intromittent organs of male squamates ( snakes, lizards and worm lizards). Hemipenes are usually held inverted within the body, and are everted for reproduction via erectile tissue, much like ...
. He added New Guinea members of ''Liasis'' and ''Morelia'' to the ''reticulatus'' group. American zoologist
Arnold G. Kluge
Arnold G. Kluge (born 1935) is professor emeritus of zoology and curator emeritus of amphibians and reptiles at the University of Michigan, Museum of Zoology. Kluge authored over 140 journal articles. He served as past president of the Willi Henn ...
performed a cladistics analysis on morphological characters and recovered a ''reticulatus'' lineage as a sister to the genus ''Python''; hence not requiring a new generic name in 1993. In a 2004 genetics study using
cytochrome b
Cytochrome b within both molecular and cell biology, is a protein found in the mitochondria of eukaryotic cells. It functions as part of the electron transport chain and is the main subunit of transmembrane cytochrome bc1 and b6f complexes.
...
DNA, Robin Lawson and colleagues recovered the reticulated python as a sister to the Australo-Papuan pythons, rather than ''Python molurus'' and its relatives.
Raymond Hoser
Raymond Terrence Hoser (born 1962) is an Australian snake-catcher and author. Since 1976, he has written books and articles about official corruption in Australia. He has also written works on Australian frogs and reptiles. Hoser's work on herpe ...
erected the genus ''Broghammerus'' for the reticulated python in 2004, naming it after German snake expert Stefan Broghammer, on the basis of dorsal patterns distinct from those of the genus ''Python'', and a dark mid-dorsal line from the rear to the front of the head, and red or orange (rather than brown) iris colour.
In 2008, Lesley Rawlings and colleagues reanalysed Kruge's morphological data and combined them with genetic material, and found the reticulated clade to be an offshoot of the Australo-Papuan lineage, as well. They adopted and redefined the genus name ''Broghammerus''.
Reynolds and colleagues also confirmed the clade's place as a sister to the Australo-Papuan pythons and coined the name ''Malayopython'', stating that the name ''Broghammerus'' was "invalid" due to it being "non-peer reviewed writing that included no formal data or analyses".
Reynolds and colleagues cite Kaiser and colleagues who state that, pending "suitable action" from the
International Commission on Zoological Nomenclature
The International Commission on Zoological Nomenclature (ICZN) is an organization dedicated to "achieving stability and sense in the scientific naming of animals". Founded in 1895, it currently comprises 26 commissioners from 20 countries.
Orga ...
(ICZN), the name ''Python'' should be used in preference of ''Broghammerus.'' In 2021, the ICZN reported that it found no basis under the provisions of the Code for regarding
oser's journalas being "unpublished" (i.e. invalid).
[ICZN. 2021. Opinion 2468 (Case 3601) - Spracklandus Hoser, 2009 (Reptilia, Serpentes, Elapidae) and Australasian Journal of Herpetology issues 1-24: confirmation of availability declined; Appendix A (Code of Ethics): not adopted as a formal criterion for ruling on cases. The Bulletin of Zoological Nomenclature 78:42–45. ]
Species
References
{{Taxonbar, from=Q20052394
Snake genera
Pythonidae