''Magericyon'' is an extinct genus of
amphicyonid
Amphicyonidae is an extinct family of terrestrial carnivorans belonging to the suborder Caniformia. They first appeared in North America in the middle Eocene (around 45 mya), spread to Europe by the late Eocene (35 mya), and further spread to Asi ...
("bear-dog") that lived during the
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
10-9 Ma (
Vallesian
The Vallesian age is a period of geologic time (11.6–9.0 Ma) within the Miocene used more specifically with European Land Mammal Ages. It precedes the Turolian age and follows the Astaracian age. The so-called Vallesian Crisis resulted in th ...
Age) in what is now
Spain
Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
.
Description
The body of this animal was vaguely similar to that of a particularly robust, large
felid
Felidae ( ) is the Family (biology), family of mammals in the Order (biology), order Carnivora colloquially referred to as cats. A member of this family is also called a felid ( ).
The 41 extant taxon, extant Felidae species exhibit the gre ...
, but the skull resembles that of a
canid
Canidae (; from Latin, ''canis'', "dog") is a family (biology), biological family of caniform carnivorans, constituting a clade. A member of this family is also called a canid (). The family includes three subfamily, subfamilies: the Caninae, a ...
or an
ursid, like that of many amphicyonids. Unlike most other amphicyonids, ''Magericyon'' had teeth associated with those of a
hypercarnivore
A hypercarnivore is an animal that has a diet that is more than 70% meat, either via active predation or by scavenging. The remaining non-meat diet may consist of non-animal foods such as fungi, fruits or other plant material. Some extant exampl ...
, with laterally flattened canines, the third premolar having a single root, the absence of second premolars, and a metaconid on its lower molars with a reduction in the second upper molar. The scapula and the front leg showed primitive features such as an
acromion
In human anatomy, the acromion (from Greek: ''akros'', "highest", ''ōmos'', "shoulder", : acromia) or summit of the shoulder is a bony process on the scapula (shoulder blade). Together with the coracoid process, it extends laterally over the sh ...
in the shoulder with a reduced caudoventral projection and post scapular pit.
[Peigné, S., Salesa, M. J., Antón, M. & Morales, J., 2008: A new amphicyonine (Carnivora: Amphicyonidae) from the upper Miocene of Batallones-1, Madrid, Spain. Palaeontology: Vol. 51, #4, pp. 943][Siliceo, G., Salesa, M. J., Antón, M., Pastor, J. F., Morales, J. 2015. Comparative Anatomy of the Shoulder Region in the Late Miocene Amphicyonid Magericyon anceps (Carnivora): Functional and Paleoecological Inferences. Journal of Mammalian Evolution, 22: 243–258.] ''Magericyon'' was roughly equivalent to a large leopard in size, weighing around .
Classification
''Magericyon'' was described for the first time in 2008, based on fossils found in
Cerro de los Batallones in Spain. The type species is ''Magericyon anceps'', but a second species has also been attributed to the genus as ''M. castellanus'', described in 1981 and initially attributed to the genus ''Amphicyon''. ''Magericyon'' is part of the family Amphicyonidae, a group of very common carnivores ranging from the Eocene to the Miocene, and which occupied many different ecological niches. ''Magericyon'' is the last amphicyonid known from Western Europe. Evidence also indicates that ''Magericyon'' was closely related to ''
Amphicyon
''Amphicyon'' is an extinct genus of large carnivorans belonging to the family Amphicyonidae (known colloquially as "bear-dogs"), subfamily Amphicyoninae, from the Miocene epoch. Members of this family received their vernacular name for possessin ...
''.
Paleobiology
''Magericyon'' occupied a different ecological niche than other amphicyonids, such as the larger ''Amphicyon'' and ''
Ysengrinia
''Ysengrinia'' is an extinct genus of carnivoran in the family Bear dog, Amphicyonidae, that lived during the Chattian, Late Oligocene to Early Miocene. Fossil remains have been discovered in Western Europe, the United States and possibly China. ...
'' (which had lifestyles more akin to bears) or ''
Daphoenodon
''Daphoenodon'' is an extinct genus of Terrestrial animal, terrestrial carnivore, which lived in the early Miocene and belonged to the family Amphicyonidae ("bear dogs") of the suborder Caniformia. The species of ''Daphoenodon'' are characteri ...
'' and ''
Temnocyon
''Temnocyon'' is an extinct genus of amphicyonids endemic to North America. It lived from the Oligocene to Early Miocene approximately 30.8—20.4 mya, existing for about .
The first fossils are recorded in North America at Logan Butte in the ...
'' of North America, which were more capable runners. ''Magericyon'' probably lived in a similar manner to that of modern felines, being an ambush hunter of large prey.
Studies by Gemo Siliceo ''et al'' also revealed that ''Magericyon'' had powerful jaw and neck muscles that helped to stabilize its head and jaws during a bite. ''Magericyon'' was particularly adept at side-to-side movements and rotations of the head. This feature allowed ''Magericyon'' to swiftly and efficiently process the meat on a carcass, allowing the animal to consume sufficient amounts of flesh before scavengers could take over its kill.
Paleoecology
As a carnivore at Cerro de los Battalones, ''Magericyon'' shared the apex predator position with two saber-toothed cat species, the leopard-sized ''
Promegantereon ogygia'' and the tiger-sized ''
Machairodus aphanistus
''Machairodus'' (from , 'knife' and 'tooth') is a genus of large machairodont or ''saber-toothed cat'' that lived in Africa and Eurasia during the Middle to Late Miocene, from 12.5 million to 8.7 million years ago. It is the animal from which ...
.'' Evidence indicates that the large carnivores may have co-existed using
niche partitioning
In ecology, a niche is the match of a species to a specific environmental condition.
Three variants of ecological niche are described by
It describes how an organism or population responds to the distribution of resources and competitors (for e ...
''.'' A carbon-13 bone analysis matched isotope profiles in prey species with their predators, showing both cats hunted prey that typically live in closed woodland habitats, such as pigs of the genus ''
Microstonyx
''Microstonyx'' was an extinct genus of suid that existed during the Miocene in Asia and Europe.
Geographic range
Fossils of the species ''M. major'' have been found in Spain, North Macedonia, Turkiye, and China
China, officially the ...
'' and perhaps occasionally young of the "tetralophodont gomphothere" ''
Tetralophodon
''Tetralophodon'' ("four-ridged tooth") is an extinct genus of "tetralophodont gomphothere" belonging to the superfamily Elephantoidea, known from the Miocene of Afro-Eurasia.
Taxonomy and evolution
The genus ''Tetralophodon'' (meaning "four-rid ...
''. ''Magericyon'' ate medium-sized prey that live in more open habitats, with the antelope ''Austroportax'' being an important food source and
hipparionine horses also present in the diet. Because the site attracted all three species as a
predator trap, ''Magericyon'' may also have taken carrion or injured animals of various kinds, though its teeth show it was specialized as a hypercarnivore without the bone-cracking adaptations of many other bear-dogs.
''Magericyon'' may have competed at times with large omnivorous bears such as ''
Agriotherium
''Agriotherium'' is an extinct genus of bears whose fossils are found in Miocene through Pleistocene-aged strata of North America, Eurasia, and Africa. The earliest species, ''A.'' ''aecuatorialis'' evolved during the early Late Miocene, around ...
'' and ''
Indarctos
''Indarctos'' is an extinct genus of bear, present in Africa, North America, and Eurasia during the Miocene. It was present from ~11.1 to 5.3 Annum, Ma, existing for approximately .
The oldest member is from Arizona (~11.1—7.7 Ma) and youngest ...
'', but these genera fed more on plant material. As well as ''Austroportax'' and ''
Hippotherium
''Hippotherium'' is an Extinction, extinct genus of Equidae, horse that lived during the Miocene through Pliocene ~13.65—6.7 Mya (unit), Mya, existing for .
The last known surviving ''Hippotherium'' was ''H. malpassii'', found in Italy.
Spe ...
,'' its prey could have included young of the hornless rhinoceros ''
Aceratherium'' and possibly the calves of the large
sivatherine giraffes and
boselaphine antelopes. Since its choice of prey suggests it lived in open country, while its legs conversely were not designed for speed, ''Magericyon'' probably would have engaged in
kleptoparasitism
Kleptoparasitism (originally spelt clepto-parasitism, meaning "parasitism by theft") is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct f ...
against any of the smaller carnivores in the region or in scavenging when the opportunity presented itself.
References
{{Taxonbar, from=Q25400575
Amphicyonidae
Miocene Amphicyonidae
Miocene mammals of Europe
Fossil taxa described in 2008
Prehistoric carnivoran genera