The

saccule The saccule (Latin: sacculus) is a bed of sensory cells in the inner ear that detects linear acceleration and head tilting in the vertical plane, and converts these vibrations into electrical impulses to be interpreted by the brain. When the he ...

is the smaller sized

vestibular The Vestibular (from , "entrance hall") is a competitive examination and is the primary and widespread entrance system used by Brazilian universities to select the students admitted. The Vestibular usually takes place from November to January, ...

sac (the utricle being the other larger size vestibular sac); it is globular in form, and lies in the recessus sphæricus near the opening of the

scala vestibuli The vestibular duct or scala vestibuli is a perilymph-filled cavity inside the cochlea of the inner ear that conducts sound vibrations to the cochlear duct. It is separated from the cochlear duct by Reissner's membrane and extends from the vesti ...

of the

cochlea The cochlea is the part of the inner ear involved in hearing. It is a spiral-shaped cavity in the bony labyrinth, in humans making 2.75 turns around its axis, the modiolus (cochlea), modiolus. A core component of the cochlea is the organ of Cort ...

. Its anterior part exhibits an oval thickening, the macula of saccule (or saccular macula), to which are distributed the saccular filaments of the

acoustic nerve The vestibulocochlear nerve or auditory vestibular nerve, also known as the eighth cranial nerve, cranial nerve VIII, or simply CN VIII, is a cranial nerve that transmits sound and equilibrium (balance) information from the inner ear to the bra ...

. The vestibule is a region of the inner ear which contains the

and the utricle, each of which contain a macula to detect

linear acceleration In mechanics, acceleration is the Rate (mathematics), rate of change of the velocity of an object with respect to time. Acceleration is one of several components of kinematics, the study of motion. Accelerations are Euclidean vector, vector ...

. Its function is to detect vertical linear acceleration. The macula of saccule lies in a nearly vertical position. It is a 2mm by 3mm patch of hair cells. Each hair cell of the macula contains 40 to 70

stereocilia Stereocilia (or stereovilli or villi) are non-motile apical cell modifications. They are distinct from cilia and microvilli, but are closely related to microvilli. They form single "finger-like" projections that may be branched, with normal cel ...

and one true

cilia The cilium (: cilia; ; in Medieval Latin and in anatomy, ''cilium'') is a short hair-like membrane protrusion from many types of eukaryotic cell. (Cilia are absent in bacteria and archaea.) The cilium has the shape of a slender threadlike proj ...

, called a

kinocilium A kinocilium is a special type of cilium on the apex of hair cells located in the sensory epithelium of the vertebrate inner ear. Contrasting with stereocilia, which are numerous, there is only one kinocilium on each hair cell. The kinocilium can ...

. A gelatinous cover called the

otolithic membrane The otolithic membrane is a fibrous structure located in the vestibular system of the inner ear. It plays a critical role in the brain's interpretation of equilibrium. The membrane serves to determine if the body or the head is tilted, in addition ...

envelops the tips of the stereocilia and kinocilium. The otolithic membrane is weighted with small densely packed

protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...

calcium carbonate Calcium carbonate is a chemical compound with the chemical formula . It is a common substance found in Rock (geology), rocks as the minerals calcite and aragonite, most notably in chalk and limestone, eggshells, gastropod shells, shellfish skel ...

granules called statoconica. The macula of the utricle is in a horizontal position and detects horizontal acceleration. The coordinated

sensory perception Perception () is the organization, identification, and interpretation of sensory information in order to represent and understand the presented information or environment. All perception involves signals that go through the nervous syste ...

of acceleration both vertically and horizontally along the vestibular nerve, allow for the perception of linear acceleration in any direction. In vertical linear acceleration, the weighted otolithic membrane lags behind the stereocilia and kinocilium. This bends the stereocilia, which is interpreted by the brain as vertical linear acceleration.

References

Vestibular system {{Anatomy-stub