''Macrovipera lebetinus'', known as the blunt-nosed viper, Lebetine viper, Levant viper,Mallow D, Ludwig D, Nilson G. 2003. ''True Vipers: Natural History and Toxinology of Old World Vipers''. Malabar, Florida: Krieger Publishing Company. 359 pp. . and by other common names, is a

viper The Viperidae (vipers) are a family of snakes found in most parts of the world, except for Antarctica, Australia, Hawaii, Madagascar, and various other isolated islands. They are venomous and have long (relative to non-vipers), hinged fangs th ...

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

found in

North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in t ...

, much of the

Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing Arabia (including the Arabian Peninsula and Bahrain), Asia Minor (Asian part of Turkey except Hatay Province), East Thrace (Europ ...

, and as far east as Kashmir. Like all other vipers, it is

venomous Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a ...

. Five subspecies are currently recognized, including the

nominate race In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all specie ...

described here.

Common names

Common names for this viper include: blunt-nosed viper, Lebetine viper, Levant viper, Levantine viper, Levantine adder,Brown JH. 1973. ''Toxicology and Pharmacology of Venoms from Poisonous Snakes''. Springfield, Illinois: Charles C. Thomas. 184 pp. LCCCN 73-229. . ''kufi'' or kufi viper (from

Arabic Arabic (, ' ; , ' or ) is a Semitic language spoken primarily across the Arab world.Semitic languages: an international handbook / edited by Stefan Weninger; in collaboration with Geoffrey Khan, Michael P. Streck, Janet C. E.Watson; Walte ...

), ''gjurza'' (from Persian), coffin snake, Levante viper, mountain viper,U.S. Navy. 1991. ''Poisonous Snakes of the World''. United States Government. New York: Dover Publications Inc. 203 pp. . ''gunas'' (from

Kashmiri Kashmiri may refer to: * People or things related to the Kashmir Valley or the broader region of Kashmir * Kashmiris, an ethnic group native to the Kashmir Valley * Kashmiri language, their language People with the name * Kashmiri Saikia Barua ...

), ''fina'' or ''kontonoura'' (the second comes from the translation of Greek Cypriot dialect for "short-tailed").

Taxonomy

This species is currently subject to review. It is likely that certain subspecies will soon be elevated to valid species status. The nominate subspecies was restricted to Cyprus in 1928 by Mertens and Müller and so does not actually occur in the

Levant The Levant () is an approximation, approximate historical geography, historical geographical term referring to a large area in the Eastern Mediterranean region of Western Asia. In its narrowest sense, which is in use today in archaeology an ...

region. The populations found in southern

Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan,; prs, امارت اسلامی افغانستان is a landlocked country located at the crossroads of Central Asia and South Asia. Referred to as the Heart of Asia, it is bord ...

and northern

India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...

are sometimes referred to as a separate subspecies: ''M. l. peilei''. These normally have semidivided supraoculars. ''Vipera Euphratica'' was originally used to refer to the populations that occur in the Euphrates river basin of Turkey, Syria and Iraq. It was synonymized with ''M. l. obtusa'' in several publications, including Joger (1984). However, Golay et al. (1993) include it in the synonymy of ''M. l. lebetina''. Obst (1983) suggested inclusion of the species in the genus ''

Daboia ''Daboia'' is a genus of venomous vipers.. Species The following four species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species ...

'' instead of ''Macrovipera''.

Subspecies

Description

This is a large snake, with females reaching 150 cm (59.1 in) in total length (body + tail) and males a little less. Sizes vary among different populations, with ''M. l. lebetina'' being somewhat smaller. The head is broad, triangular, and distinct from the neck. The snout is rounded and blunt when viewed from above, which is why it is also called the blunt-nosed viper. The

nasal Nasal is an adjective referring to the nose, part of human or animal anatomy. It may also be shorthand for the following uses in combination: * With reference to the human nose: ** Nasal administration, a method of pharmaceutical drug delivery ** ...

and

nasorostral scales In reptiles, the nasorostral is an enlarged and usually paired scale, just behind the rostral Rostral may refer to: Anatomy * Rostral (anatomical term), situated toward the oral or nasal region * Rostral bone, in ceratopsian dinosaurs * Rostral or ...

are almost completely fused into a single plate, although some variation occurs. The

dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...

are strongly keeled, except for those bordering the

ventrals In snakes, the ventral scales or gastrosteges are the enlarged and transversely elongated scales that extend down the underside of the body from the neck to the anal scale. When counting them, the first is the anteriormost ventral scale that cont ...

. ''M. l. lebetina'' usually has 146-163 ventral scales. The

anal scale Anal may refer to: Related to the anus *Related to the anus of animals: ** Anal fin, in fish anatomy ** Anal vein, in insect anatomy ** Anal scale, in reptile anatomy *Related to the human anus: ** Anal sex, a type of sexual activity involving ...

is single. The color pattern is less varied than one might expect from a species that is so widely distributed. The head is normally uniformly colored, although it can occasionally be marked with a dark V-shape. Dorsally, the ground color of the body can be gray, brown, beige, pinkish, olive, or khaki. The pattern, if present, is darker. It can be gray, bluish, rust, or brown in color, and may consist of a middorsal row or double row of large spots. When two rows are present, the spots may alternate or oppose, which can produce anything from a saddled to a continuous zigzag pattern. The spots are usually brown, dark gray, or black, but are sometimes red, brick, yellow, or olive in color. Males are usually 3 and a half feet (1.1 m) in total length, while females may attain a total length of 5 feet (1.5 m).

Habitat

It may be found under short trees with thick shade if the outside temperature is greater than 45° Celsius.

Geographic range

The blunt-nosed viper can be found in

Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , religi ...

Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...

Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is ...

Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...

, Syria,

Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Ri ...

Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...

Lebanon Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to Lebanon–Syria border, the north and east and Israel to Blue ...

Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...

Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkm ...

, Russian Caucasia,

Armenia Armenia (), , group=pron officially the Republic of Armenia,, is a landlocked country in the Armenian Highlands of Western Asia.The UNbr>classification of world regions places Armenia in Western Asia; the CIA World Factbook , , and ...

Georgia Georgia most commonly refers to: * Georgia (country), a country in the Caucasus region of Eurasia * Georgia (U.S. state), a state in the Southeast United States Georgia may also refer to: Places Historical states and entities * Related to t ...

Azerbaijan Azerbaijan (, ; az, Azərbaycan ), officially the Republic of Azerbaijan, , also sometimes officially called the Azerbaijan Republic is a transcontinental country located at the boundary of Eastern Europe and Western Asia. It is a part of th ...

Turkmenistan Turkmenistan ( or ; tk, Türkmenistan / Түркменистан, ) is a country located in Central Asia, bordered by Kazakhstan to the northwest, Uzbekistan to the north, east and northeast, Afghanistan to the southeast, Iran to the s ...

Uzbekistan Uzbekistan (, ; uz, Ozbekiston, italic=yes / , ; russian: Узбекистан), officially the Republic of Uzbekistan ( uz, Ozbekiston Respublikasi, italic=yes / ; russian: Республика Узбекистан), is a doubly landlocked co ...

Kazakhstan Kazakhstan, officially the Republic of Kazakhstan, is a transcontinental country located mainly in Central Asia and partly in Eastern Europe. It borders Russia Russia (, , ), or the Russian Federation, is a transcontinental coun ...

Tajikistan Tajikistan (, ; tg, Тоҷикистон, Tojikiston; russian: Таджикистан, Tadzhikistan), officially the Republic of Tajikistan ( tg, Ҷумҳурии Тоҷикистон, Jumhurii Tojikiston), is a landlocked country in Centr ...

Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...

, and Kashmir,

. Scortecci (1929) also reported this species from

Yemen Yemen (; ar, ٱلْيَمَن, al-Yaman), officially the Republic of Yemen,, ) is a country in Western Asia. It is situated on the southern end of the Arabian Peninsula, and borders Saudi Arabia to the north and Oman to the northeast an ...

. The

type locality Type locality may refer to: * Type locality (biology) * Type locality (geology) See also * Local (disambiguation) * Locality (disambiguation) {{disambiguation ...

originally given was ''"Oriente"''.

Mertens __NOTOC__ Mertens () is a surname of Flemish Origin, meaning "son of Merten" (Martin). It is the fifth most common name in Belgium with 18,518 people in 2008. Geographical distribution As of 2014, 43.4% of all known bearers of the surname ''Merte ...

and L. Müller (1928) suggested restricting the range to ''"Cypern"'' (= Cyprus).

Conservation status

Lebetine vipers are endangered. The species is listed as strictly protected (Appendix II) under the

Berne Convention The Berne Convention for the Protection of Literary and Artistic Works, usually known as the Berne Convention, was an international assembly held in 1886 in the Swiss city of Bern by ten European countries with the goal to agree on a set of le ...

.Convention on the Conservation of European Wildlife and Natural Habitats, Appendix II
a
Council of Europe
Accessed 9 October 2006.