''Macroprotodon cucullatus'', commonly known as the false smooth snake, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of mildly

venom Venom or zootoxin is a type of toxin produced by an animal that is actively delivered through a wound by means of a bite, sting, or similar action. The toxin is delivered through a specially evolved ''venom apparatus'', such as fangs or a sti ...

ous

colubrid Colubridae (, commonly known as colubrids , from , 'snake') is a family of snakes. With 249 genera, it is the largest snake family. The earliest fossil species of the family date back to the Late Eocene epoch, with earlier origins suspected. Colu ...

snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the Mediterranean Basin.

Geographic distribution

''M. cucullatus'' is found in

Algeria Algeria, officially the People's Democratic Republic of Algeria, is a country in the Maghreb region of North Africa. It is bordered to Algeria–Tunisia border, the northeast by Tunisia; to Algeria–Libya border, the east by Libya; to Alger ...

Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...

Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...

Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...

Libya Libya, officially the State of Libya, is a country in the Maghreb region of North Africa. It borders the Mediterranean Sea to the north, Egypt to Egypt–Libya border, the east, Sudan to Libya–Sudan border, the southeast, Chad to Chad–L ...

Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...

Palestinian Territories The occupied Palestinian territories, also referred to as the Palestinian territories, consist of the West Bank (including East Jerusalem) and the Gaza Strip—two regions of the former Mandate for Palestine, British Mandate for Palestine ...

Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...

Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...

, and

Tunisia Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...

Description

''Macroprotodon cucullatus'' is a small snake, usually not exceeding in total length (including tail). As the common name implies, the

dorsal scales In snakes, the dorsal scales are the longitudinal series of plates that encircle the body, but do not include the ventral scales. Campbell JA, Lamar WW (2004). ''The Venomous Reptiles of the Western Hemisphere''. Ithaca and London: Comstock Publis ...

are smooth, and are arranged in 19-23 rows.

Dorsally Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provi ...

it is tan or gray, with small brown spots, or with darker and lighter streaks. Ventrally it is yellow to coral-red, uniform or spotted with black, the spots sometimes confluent at the midline. It has a blackish collar which may extend to the top of the head. There is usually a dark streak from the nostril to the corner of the mouth, passing under or through the eye. The eye is rather small, and the pupil, when seen contracted in bright light, is oval. The snout is flattened, and the

rostral Rostral may refer to: Anatomy * Rostral (anatomical term), situated toward the oral or nasal region * Rostral bone, in ceratopsian dinosaurs * Rostral organ, of certain fish * Rostral scale The rostral scale, or rostral, in snakes and other sca ...

is wide and low, barely visible from above. The 6th upper labial extends upward and contacts the parietal. There is usually one anterior temporal. Arnold, E.N., and J.A. Burton (1978). ''A Field Guide to the Reptiles and Amphibians of Britain and Europe''. London: Collins. 272 pp. . (''Macroprotodon cucullatus'', pp. 206-207 + Plate 38 + Map 119).

Habitat

The natural

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s of ''M. cucullatus'' are temperate

forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...

s, temperate

shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...

, subtropical or tropical dry shrubland, Mediterranean-type shrubby vegetation, freshwater spring, rocky areas, sandy shores,

arable land Arable land (from the , "able to be ploughed") is any land capable of being ploughed and used to grow crops.''Oxford English Dictionary'', "arable, ''adj''. and ''n.''" Oxford University Press (Oxford), 2013. Alternatively, for the purposes of a ...

, pastureland,

plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...

s, rural gardens, and

urban area An urban area is a human settlement with a high population density and an infrastructure of built environment. Urban areas originate through urbanization, and researchers categorize them as cities, towns, conurbations or suburbs. In urbani ...

Behaviour

The false smooth snake is nocturnal in some parts of its range but mainly diurnal in the Balearic Islands.

Diet

''M. cucullatus'' preys upon small

lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...

s such as

gecko Geckos are small, mostly carnivorous lizards that have a wide distribution, found on every continent except Antarctica. Belonging to the infraorder Gekkota, geckos are found in warm climates. They range from . Geckos are unique among lizards ...

s and lizards of the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

Lacerta Lacerta is one of the 88 modern constellations defined by the International Astronomical Union. Its name is Latin for lizard. A small, faint constellation, it was defined in 1687 by the astronomer Johannes Hevelius. Its brightest stars form a "W" ...

''. Small mammals and nestling birds are also eaten.

Reproduction

Females of ''M. cucullatus'' may breed in alternate years. Two to six eggs are laid in a damp spot under a stone, buried in the soil or hidden in dense vegetation. They hatch in about eight weeks into juveniles with a snout-to-vent length of .

Venom

''M. cucullatus'' possesses a mild venom, which is delivered by means of enlarged grooved teeth in the upper jaw. Although the venom is effective on lizards, this snake is not harmful to humans due to its small size.

Conservation status

The

IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...

has listed the false smooth snake as being of "

Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...

". This is because it has a wide distribution and a large population, seems to be tolerant of some habitat modification and its population is unlikely to be declining fast enough to qualify it for listing in a more threatened category.

References

External links

*Corti, C.; Mellado, V.P.; Geniez, P.; El Din, S.B.; Martinez-Solano, I; Sindaco, R.; Romano, A. (2009). IUCN (International Union for Conservation of Nature and Natural Resources) Red List Assessment, Version 3.1. http://eol.org/data_objects/18923870. Accessed October 2013.
''Macroprotodon cucullatus'' (Geoffroy Saint-Hilaire, 1827)
. {{Taxonbar, from=Q1728800 Macroprotodon Snakes of Africa Snakes of Asia Reptiles of Europe Fauna of the Middle East Reptiles of North Africa Reptiles described in 1827 Taxa named by Isidore Geoffroy Saint-Hilaire Taxonomy articles created by Polbot