Macronectes Halli
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The northern giant petrel (''Macronectes halli''), also known as Hall's giant petrel, is a large, predatory
seabird Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...
of the southern oceans. Its range overlaps broadly with the range of the related
southern giant petrel The southern giant petrel (''Macronectes giganteus''), also known as the Antarctic giant petrel, giant fulmar, stinker, and stinkpot, is a large seabird of the southern oceans. Its distribution overlaps broadly with the similar northern giant pe ...
(''Macronectes giganteus''), albeit slightly further to the north.


Taxonomy

The northern giant petrel was formally described in 1912 by the Australian born ornithologist
Gregory Mathews Gregory Macalister Mathews CBE FRSE FZS FLS (10 September 1876 – 27 March 1949) was an Australian-born amateur ornithologist who spent most of his later life in England. Life He was born in Biamble in New South Wales the son of Robert H. M ...
as a
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of the
southern giant petrel The southern giant petrel (''Macronectes giganteus''), also known as the Antarctic giant petrel, giant fulmar, stinker, and stinkpot, is a large seabird of the southern oceans. Its distribution overlaps broadly with the similar northern giant pe ...
with the
trinomial name In biology, trinomial nomenclature is the system of names for taxa below the rank of species. These names have three parts. The usage is different in zoology and botany. In zoology In zoological nomenclature, a trinomen (), trinominal name, or t ...
''Macronectes giganteus halli''. The specific epithet ''halli'' was chosen to honour the Australian ornithologist Robert Hall who had described the birds breeding on the
Kerguelen Islands The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the subantarctic, sub-Antarctic region. They are among the Extremes on Earth#Remoteness, most i ...
. The northern giant petrel is now considered to be a separate species and has the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Macronectes halli''. It is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised. The genus name combines the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''makros'' meaning "great" and ''nēktēs'' meaning "swimmer". The name "petrel" refers to the Biblical account of
Saint Peter Saint Peter (born Shimon Bar Yonah; 1 BC – AD 64/68), also known as Peter the Apostle, Simon Peter, Simeon, Simon, or Cephas, was one of the Twelve Apostles of Jesus and one of the first leaders of the Jewish Christian#Jerusalem ekklēsia, e ...
walking on water, referring to the way these birds run on top of the water as they are getting airborne. The two giant petrel species make up the genus '' Macronectes''. They belong to the
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
Procellariiformes Procellariiformes is an order (biology), order of seabirds that comprises four family (biology), families: the albatrosses, the Procellariidae, petrels and shearwaters, and two families of storm petrels. Formerly called Tubinares and still call ...
, the tube-nosed seabirds or petrels. All tube-noses have tubular nostrils, and all those in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Procellariidae The family (biology), family Procellariidae is a group of seabirds that comprises the fulmarine petrels, the gadfly petrels, the diving petrels, the prion (bird), prions, and the shearwaters. This family is part of the bird order (biology), orde ...
, the true petrels, have their nostrils united along the top of the bill. Procellariform birds have between seven and nine distinct horny plates for their bill, and in petrels one of these plates forms the hooked portion of their upper bill called the maxillary unguis. They produce a
stomach oil Stomach oil is the light oil composed of neutral dietary lipids found in the proventriculus (fore-gut) of birds in the order Procellariiformes. All albatrosses, procellarids (gadfly petrels and shearwaters) and northern and austral storm ...
made up of
wax ester A wax ester (WE) is an ester of a fatty acid and a fatty alcohol. Wax esters are the main components of three commercially important waxes: carnauba wax, candelilla wax, and beeswax.. Wax esters are formed by combining one fatty acid with one ...
s and
triglycerides A triglyceride (from ''wikt:tri-#Prefix, tri-'' and ''glyceride''; also TG, triacylglycerol, TAG, or triacylglyceride) is an ester derived from glycerol and three fatty acids. Triglycerides are the main constituents of body fat in humans and oth ...
that is stored in the
proventriculus The proventriculus is part of the digestive system of birds.Encarta World English Dictionary orth American Edition(2007). ''Proventriculus''. Source: (accessed: December 18, 2007) An analogous organ exists in invertebrates and insects. Birds Th ...
. This can be sprayed out of their mouths as a defence against predators as well as an energy rich food source for chicks and for the adults during their long flights. Finally, they also have a
salt gland The salt gland is an organ (anatomy), organ for excreting excess salt (chemistry), salts. It is found in the cartilaginous fishes subclass elasmobranchii (sharks, rays, and skates), seabirds, and some reptiles. Salt glands can be found in the r ...
that is situated above the nasal passage and helps desalinate their bodies, due to the high amount of sea water that they imbibe. It excretes a concentrated saline solution from their nostrils.


Description

The northern giant petrel averages in length, with a range of , possessing a wingspan of . Size is somewhat variable and this species is broadly the same size as its southern sister species. The largest-bodied colony is in the
South Georgia Islands South Georgia and the South Sandwich Islands (SGSSI) is a British Overseas Territory in the southern Atlantic Ocean. It is a remote and inhospitable collection of islands, consisting of South Georgia and a chain of smaller islands known as the S ...
, where 56 males averaged and 43 females average .''CRC Handbook of Avian Body Masses, 2nd Edition'' by John B. Dunning Jr. (Editor). CRC Press (2008), . The smallest-bodied are on the
Chatham Islands The Chatham Islands ( ; Moriori language, Moriori: , 'Misty Sun'; ) are an archipelago in the Pacific Ocean about east of New Zealand's South Island, administered as part of New Zealand, and consisting of about 10 islands within an approxima ...
, where 19 males averaged and 21 females averaged .Carlos, C. J., & Voisin, J. F. (2008). ''Identifying giant petrels, Macronectes giganteus and M. halli, in the field and in the hand''. Publishing Editor, 1. Overall, weight for the species can range from . Its
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
consists of grey-brown body with lighter coloured forehead, sides of face, and chin. Its bill is between long, slightly longer on average than the southern giant petrel, and is pinkish yellow with a brown tip. Its eyes are grey. The tarsus of the northern giant petrel is slightly longer on average than the southern species, but the southern has longer wings on average. The juvenile ''Macronectes halli'' is entirely dark brown and lightens as it ages. It can be differentiated from ''Macronectes giganteus'' by the top of the bill, which on the southern species is green.


Distribution and habitat

The northern giant petrel is
pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...
and is found throughout the
Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the world ocean, generally taken to be south of 60th parallel south, 60° S latitude and encircling Antarctica. With a size of , it is the seco ...
north of the Antarctic Convergence Zone, and north through Chile,
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
, South Africa, New Zealand, and half of Australia. Over 4,500 pairs nest on islands in the
South Georgia South Georgia is an island in the Atlantic Ocean, South Atlantic Ocean that is part of the British Overseas Territories, British Overseas Territory of South Georgia and the South Sandwich Islands. It lies around east of the Falkland Islands. ...
group. They also nest on some of the
Chatham Islands The Chatham Islands ( ; Moriori language, Moriori: , 'Misty Sun'; ) are an archipelago in the Pacific Ocean about east of New Zealand's South Island, administered as part of New Zealand, and consisting of about 10 islands within an approxima ...
,
Kerguelen Islands The Kerguelen Islands ( or ; in French commonly ' but officially ', ), also known as the Desolation Islands (' in French), are a group of islands in the subantarctic, sub-Antarctic region. They are among the Extremes on Earth#Remoteness, most i ...
,
Crozet Islands The Crozet Islands (; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarctic Lands. History ...
,
Macquarie Island Macquarie Island is a subantarctic island in the south-western Pacific Ocean, about halfway between New Zealand and Antarctica. It has been governed as a part of Tasmania, Australia, since 1880. It became a Protected areas of Tasmania, Tasmania ...
and others. Their overall range is .


Behaviour


Feeding

The northern giant petrel feeds mainly on
carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
(dead penguins and
pinniped Pinnipeds (pronounced ), commonly known as seals, are a widely range (biology), distributed and diverse clade of carnivorous, fin-footed, semiaquatic, mostly marine mammals. They comprise the extant taxon, extant families Odobenidae (whose onl ...
s), as well as fish,
krill Krill ''(Euphausiids)'' (: krill) are small and exclusively marine crustaceans of the order (biology), order Euphausiacea, found in all of the world's oceans. The name "krill" comes from the Norwegian language, Norwegian word ', meaning "small ...
,
squid A squid (: squid) is a mollusc with an elongated soft body, large eyes, eight cephalopod limb, arms, and two tentacles in the orders Myopsida, Oegopsida, and Bathyteuthida (though many other molluscs within the broader Neocoleoidea are also ...
, and other
cephalopods A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...
. They will follow
fishing boats A fishing vessel is a boat or ship used to fishing, catch fish and other valuable nektonic aquatic animals (e.g. shrimps/prawns, krills, coleoids, etc.) in the sea, lake or river. Humans have used different kinds of surface vessels in commercial ...
and
cruise ships Cruise ships are large passenger ships used mainly for vacationing. Unlike ocean liners, which are used for transport, cruise ships typically embark on round-trip voyages to various ports of call, where passengers may go on tours known as " ...
, eating any discarded fish and waste from the ships. During the breeding season, males eat more carrion than females, with the females feeding on
pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...
sources. They are extremely aggressive and will kill other seabirds (mostly penguin chicks, sick or injured adult penguins and the chicks of other seabirds), even those as large as an
albatross Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Paci ...
.BirdLife International (2009) Northern giant petrels forage in similar locations to southern giant petrels, but at different times due to the earlier breeding season of northern giant petrels. It is thought this temporal segregation in habitat-use reduces interspecific competition, whilst sexual segregation, due to females making more pelagic trips than males, reduces intraspecific competition.


Breeding

Birds start breeding at an average age of ten years, and breed on islands in
colonies A colony is a territory subject to a form of foreign rule, which rules the territory and its indigenous peoples separated from the foreign rulers, the colonizer, and their '' metropole'' (or "mother country"). This separated rule was often or ...
, which they share with the
southern giant petrel The southern giant petrel (''Macronectes giganteus''), also known as the Antarctic giant petrel, giant fulmar, stinker, and stinkpot, is a large seabird of the southern oceans. Its distribution overlaps broadly with the similar northern giant pe ...
. They breed six weeks earlier than their counterparts.


Conservation

In a 2001 estimate, this species had between 17,000 and 21,000 mature birds. This number has been increasing over the last two decades, after being expected to decrease. The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
thus downgraded them from
near threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
to
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
. Recent surveys have shown that most locations are increasing in numbers, except for the
Crozet Islands The Crozet Islands (; or, officially, ''Archipel Crozet'') are a sub-Antarctic archipelago of small islands in the southern Indian Ocean. They form one of the five administrative districts of the French Southern and Antarctic Lands. History ...
. This is probably due to increases in fish waste, better control on
longline fishing Longline fishing, or longlining, is a commercial fishing angling technique that uses a long ''main line'' with baited hooks attached at intervals via short branch lines called ''snoods'' or ''gangions''.carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
from fur seals. Currently, this bird is listed on CMS Appendix II and ACAP Annex 1. Future plans are to maintain surveys and counting of individuals, research movement and migration, and continue lowering the
bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...
deaths by current means and if needed newer measures through
CCAMLR The Convention on the Conservation of Antarctic Marine Living Resources, also known as the Commission for the Conservation of Antarctic Marine Living Resources, and CCAMLR, is part of the Antarctic Treaty System. The convention was opened for s ...
,
CMS CMS may refer to: Computing * Call management system * CMS-2, a programming language implemented for and used by the United States Navy * Code Morphing Software, a technology used by Transmeta * Collection management system for a museum coll ...
, and
FAO The Food and Agriculture Organization of the United Nations; . (FAO) is a List of specialized agencies of the United Nations, specialized agency of the United Nations that leads international efforts to defeat hunger and improve nutrition ...
.


References


Sources

* * * * *


External links


Species factsheet
by BirdLife International
Photos
by Christopher Taylor Nature Photography * Southern and Northern Giant Petrels â€
Species text in The Atlas of Southern African Birds
* Northern giant petrel discussed in RNZ ''Critter of the Week''
7 July 2017
{{Taxonbar, from=Q276212
northern giant petrel The northern giant petrel (''Macronectes halli''), also known as Hall's giant petrel, is a large, predatory seabird of the southern oceans. Its range overlaps broadly with the range of the related southern giant petrel (''Macronectes giganteus'') ...
Birds of Uruguay Birds of Patagonia Birds of Southern Africa Birds of New South Wales Birds of South Australia Birds of Victoria (state) Birds of Western Australia Birds of Tasmania Birds of New Zealand Birds of subantarctic islands
northern giant petrel The northern giant petrel (''Macronectes halli''), also known as Hall's giant petrel, is a large, predatory seabird of the southern oceans. Its range overlaps broadly with the range of the related southern giant petrel (''Macronectes giganteus'') ...
Fauna of Macquarie Island