Macrobaenidae is an extinct family of

turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...

s, known from the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

Paleogene The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...

Laurasia Laurasia () was the more northern of two large landmasses that formed part of the Pangaea supercontinent from around ( Mya), the other being Gondwana. It separated from Gondwana (beginning in the late Triassic period) during the breakup of Pa ...

. Their relationships to other turtles and whether they form a monophlyletic group are controversial. They are typically interpreted as stem or

crown group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor ...

cryptodires, but some more recent analyses have found them to lie outside

Testudines. Macrobaenids can be distinguished from other testudinatans by the presence of a carotid fenestra, cruciform plastron with strap-like epiplastra, and a lack of extragulars.

Genera

*'' Anatolemys'' Central Asia, Late Cretaceous ( Khodzhakul Formation, Uzbekistan,

Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...

Bissekty Formation The Bissekty Formation (sometimes referred to as Bissekt) is a formation (geology), geologic formation and Lagerstätte which outcrop, crops out in the Kyzyl Kum desert of Uzbekistan, and dates to the Late Cretaceous Period (geology), Period. Laid ...

, Uzbekistan,

Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...

, Bostobe Formation, Kazakhstan,

Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...

Yalovach Formation, Tajikistan, Santonian) *'' Appalachemys'' Mooreville Chalk, Alabama, Late Cretaceous (Santonian–

Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...

) *'' Asiachelys'' Khulsangol Formation, Mongolia, Early Cretaceous (

Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...

) *'' Aurorachelys''

Kanguk Formation The Kanguk Formation is a geological formation in the Northwest Territories and Nunavut, Canada whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B ...

, Canada, Turonian *'' Changmachelys'' Xiagou Formation, China, Early Cretaceous (Aptian) *'' Kirgizemys'' Murtoi Formation, Khilok Formation, Ilek Formation, Russia, Early Cretaceous (

Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...

Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...

), Alamyshik Formation, Kyrgyzstan, Albian, Geoncheonri Formation, South Korea, Albian Khodzhakul Formation, Uzbekistan, Cenomanian *'' Judithemys'' North America, Late Cretaceous–

Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...

*'' Gallica''

Paris Basin The Paris Basin () is one of the major geological regions of France. It developed since the Triassic over remnant uplands of the Variscan orogeny (Hercynian orogeny). The sedimentary basin, no longer a single drainage basin, is a large sag in ...

, France, Paleocene *'' Macrobaena'' Naran Bulak Formation, Mongolia, Paleocene *'' Osteopygis'' Late Cretaceous-Paleocene, North America *'' Oxemys'' Khodzhakul Formation, Uzbekistan, Cenomanian *'' Yakemys'' Phu Kradung Formation, Thailand, Early Cretaceous (

Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...

) '' Hongkongochelys'' from the Middle-Late Jurassic of China has sometimes been attributed to the family, but other times has been attriuted to Sinemydidae, a group which has an unresolved relationship with Macrobaenidae.

References

{{Testudines Extinct turtles †