Mackerel is a common name applied to a number of different species of

pelagic fish Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that live on or near the bottom, and reef fish that are associated with coral reefs. ...

, mostly from the family

Scombridae The mackerel, tuna, and bonito family, Scombridae, includes many of the most important and familiar food fishes. The family consists of 51 species in 15 genera and two subfamilies. All species are in the subfamily Scombrinae, except the but ...

. They are found in both temperate and tropical seas, mostly living along the coast or offshore in the oceanic environment. Mackerel species typically have deeply forked tails and vertical "

tiger The tiger (''Panthera tigris'') is a large Felidae, cat and a member of the genus ''Panthera'' native to Asia. It has a powerful, muscular body with a large head and paws, a long tail and orange fur with black, mostly vertical stripes. It is ...

-like" stripes on their backs with an

iridescent Iridescence (also known as goniochromism) is the phenomenon of certain surfaces that appear gradually to change colour as the angle of view or the angle of illumination changes. Iridescence is caused by wave interference of light in microstruc ...

green-blue quality. Many are restricted in their distribution ranges and live in separate populations or fish stocks based on geography. Some stocks migrate in large

schools A school is the educational institution (and, in the case of in-person learning, the building) designed to provide learning environments for the teaching of students, usually under the direction of teachers. Most countries have systems of ...

along the coast to suitable spawning grounds, where they spawn in fairly shallow waters. After spawning they return the way they came in smaller schools to suitable feeding grounds, often near an area of

upwelling Upwelling is an physical oceanography, oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted sur ...

. From there they may move offshore into deeper waters and spend the winter in relative inactivity. Other stocks migrate across oceans. Smaller mackerel are

forage fish Forage fish, also called prey fish or bait fish, are small pelagic fish that feed on planktons (i.e. planktivores) and other small aquatic organisms (e.g. krill). They are in turn preyed upon by various predators including larger fish, seabirds ...

for larger predators, including larger mackerel and

Atlantic cod The Atlantic cod (: cod; ''Gadus morhua'') is a fish of the family Gadidae, widely consumed by humans. It is also commercially known as '' cod'' or ''codling''.whales Whales are a widely distributed and diverse group of fully Aquatic animal, aquatic placental mammal, placental marine mammals. As an informal and Colloquialism, colloquial grouping, they correspond to large members of the infraorder Cetacea ...

, dolphins, sharks, and schools of larger fish such as

tuna A tuna (: tunas or tuna) is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae ( mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bul ...

and

marlin Marlins are fish from the family Istiophoridae, which includes between 9 and 11 species, depending on the taxonomic authority. Name The family's common name is thought to derive from their resemblance to a sailor's marlinspike. Taxonomy T ...

follow mackerel schools and attack them in sophisticated and cooperative ways. Mackerel flesh is high in omega-3 oils and is intensively harvested by humans. In 2009, over 5 million tons were landed by commercial fishermen. Sport fishermen value the fighting abilities of the king mackerel.

Species

Over 30 different species, principally belonging to the family

, are commonly referred to as mackerel. The term "mackerel" is derived from

Old French Old French (, , ; ) was the language spoken in most of the northern half of France approximately between the late 8th

spawn Spawn or spawning may refer to: * Spawning, the eggs and sperm of aquatic animals Arts, entertainment and media * Spawn (character), a fictional character in the comic series of the same name and in the associated franchise ** ''Spawn: Armageddon' ...

enthusiastically in Shoal">shoals near the coast, and medieval ideas on animal procreation were creative.

Scombroid mackerels

About 21 species in the family Scombridae are commonly called mackerel. The

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

for the scombroid mackerel is the Atlantic mackerel, ''Scomber scombrus''. Until recently, Atlantic chub mackerel and Indo-Pacific chub mackerel were thought to be subspecies of the same species. In 1999, Collette established, on molecular and morphological considerations, that these are separate species. Mackerel are smaller with shorter lifecycles than their close relatives, the

, which are also members of the same family.

Scombrini, the true mackerels

The true mackerels belong to the

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...

Scombrini. The tribe consists of seven species, each belonging to one of two

genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...

: '' Scomber'' or ''

Rastrelliger ''Rastrelliger'' is a mackerel genus in the family Scombridae. The three species of ''Rastrelliger'' together with the four species of ''Scomber'' comprise the tribe (biology), tribe Scombrini, known as the "true mackerels". Species The three s ...

''.

Scomberomorini, the Spanish mackerels

The

Spanish mackerel Scomberomorini is a tribe of ray-finned, saltwater, bony fishes that is commonly known as Spanish mackerels, seerfishes, or seer fish. This tribe is a subset of the mackerel family (Scombridae), which it shares with four sister tribes, the tu ...

s belong to the tribe

Scomberomorini Scomberomorini is a tribe of ray-finned, saltwater, bony fishes that is commonly known as Spanish mackerels, seerfishes, or seer fish. This tribe is a subset of the mackerel family (Scombridae), which it shares with four sister tribes, the tu ...

, which is the "cousin tribe" of the true mackerels. This tribe consists of 21 species in all—18 of those are classified into the genus '' Scomberomorus'', two into ''

Grammatorcynus ''Grammatorcynus'' is a genus of ray-finned bony fish in the family Scombridae. This genus together with '' Acanthocybium'' and '' Scomberomorus'' are comprised by the tribe Scomberomorini, commonly known as the Spanish mackerels or seerfish ...

'', and a single species into the

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

genus ''

Acanthocybium The wahoo (''Acanthocybium solandri'') is a Scombridae, scombrid fish found Cosmopolitan distribution, worldwide in tropical and subtropical seas. In Hawaii, the wahoo is known as ono. The species is sometimes called hoo in the United States. It ...

''.

Other mackerel

In addition, a number of species with mackerel-like characteristics in the families

Carangidae The Carangidae are a family of ray-finned fish that includes the jacks, pompanos, jack mackerels, runners, trevallies, and scads. It is the largest of the six families included within the order Carangiformes. Some authorities classify it as the ...

Hexagrammidae Hexagrammidae, the greenlings, is a Family (biology), family of marine ray-finned fishes belonging to the suborder Cottoidei in the Order (biology), order Perciformes. These fishes are found in the North Pacific Ocean. Taxonomy Hexagrammidae wa ...

and Gempylidae are commonly referred to as mackerel. Some confusion had occurred between the Pacific jack mackerel (''Trachurus symmetricus'') and the heavily harvested Chilean jack mackerel (''T. murphyi''). These have been thought at times to be the same species, but are now recognized as separate species. The term "mackerel" is also used as a modifier in the common names of other fish, sometimes indicating the fish has vertical stripes similar to a scombroid mackerel: * Mackerel icefish—''Champsocephalus gunnari'' * Mackerel pike—''Cololabis saira'' *

Mackerel shark The Lamniformes (, from Greek ''lamna'' "fish of prey") are an order of sharks commonly known as mackerel sharks (which may also refer specifically to the family Lamnidae). It includes some of the most familiar species of sharks, such as the g ...

—several species * Shortfin mako shark—''Isurus oxyrinchus'' *

Mackerel tuna ''Euthynnus affinis'', the mackerel tuna, little tuna, eastern little tuna, wavyback skipjack tuna, kawakawa, or tongkol komo is a species of ray-finned bony fish in the family (biology), family Scombridae, or mackerel family. It belongs to the ...

—''Euthynnus affinis'' * Mackerel tail goldfish—''Carassius auratus'' By extension, the term is applied also to other species such as the mackerel tabby cat, and to inanimate objects such as the altocumulus mackerel sky cloud formation.

Characteristics (Scombridae)

Most mackerel belong to the family Scombridae, which also includes

and

bonito Bonitos are a tribe of medium-sized, ray-finned, predatory fish in the family Scombridae, which it shares with the mackerel, tuna, and Spanish mackerel tribes, and also the butterfly kingfish. Also called the tribe Sardini, it consists of ...

. Generally, mackerel are much smaller and slimmer than tuna, though in other respects, they share many common characteristics. Their

scales Scale or scales may refer to: Mathematics * Scale (descriptive set theory), an object defined on a set of points * Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original * Scale factor, a number ...

, if present at all, are extremely small. Like tuna and bonito, mackerel are voracious feeders, and are swift and manoeuvrable swimmers, able to streamline themselves by retracting their fins into grooves on their bodies. Like other scombroids, they lack a

swim bladder The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ (anatomy), organ in bony fish that functions to modulate buoyancy, and thus allowing the fish to stay at desired water depth without having to maintain lift ...

, and their bodies are cylindrical with numerous finlets on the

dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...

and

ventral Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position prov ...

sides behind the

and

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

s, but unlike the deep-bodied tuna, they are slim. The

for scombroid mackerels is the Atlantic mackerel, ''Scomber scombrus''. These fish are

blue-green above with a silvery underbelly and near-vertical wavy black stripes running along their upper bodies. The prominent stripes on the back of mackerels seemingly are there to provide camouflage against broken backgrounds. That is not the case, though, because mackerel live in midwater pelagic environments which have no background. However, fish have an optokinetic reflex in their visual systems that can be sensitive to moving stripes. For fish to school efficiently, they need feedback mechanisms that help them align themselves with adjacent fish, and match their speed. The stripes on neighbouring fish provide "schooling marks", which signal changes in relative position. The

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

also helps with orderly schooling. A layer of thin, reflecting platelets is seen on some of the mackerel stripes. In 1998, E J Denton and D M Rowe argued that these platelets transmit additional information to other fish about how a given fish moves. As the orientation of the fish changes relative to another fish, the amount of light reflected to the second fish by this layer also changes. This sensitivity to orientation gives the mackerel "considerable advantages in being able to react quickly while schooling and feeding." Mackerel range in size from small

to larger

game fish Game fish, sport fish or quarry refer to popular fish species pursued by recreational fishing, recreational fishers (typically angling, anglers), and can be freshwater fish, freshwater or saltwater fish. Game fish can be fish as food, eaten aft ...

. Coastal mackerel tend to be small. The king mackerel is an example of a larger mackerel. Most fish are cold-blooded, but exceptions exist. Certain species of fish maintain elevated body temperatures.

Endothermic An endothermic process is a chemical or physical process that absorbs heat from its surroundings. In terms of thermodynamics, it is a thermodynamic process with an increase in the enthalpy (or internal energy ) of the system.Oxtoby, D. W; Gillis, ...

bony fish Osteichthyes ( ; ), also known as osteichthyans or commonly referred to as the bony fish, is a Biodiversity, diverse clade of vertebrate animals that have endoskeletons primarily composed of bone tissue. They can be contrasted with the Chondricht ...

es are all in the suborder Scombroidei and include the butterfly mackerel, a species of primitive mackerel. Mackerel are strong swimmers. Known in the latin family as "punctualis piscis" which translates to "punctual fish." This is due to its punctuality of migration during mating season as it moves from warm to cold waters. Atlantic mackerel can swim at a sustained speed of 0.98 m/sec with a burst speed of 5.5 m/sec, while chub mackerel can swim at a sustained speed of 0.92 m/sec with a burst speed of 2.25 m/sec. The

Wahoo The wahoo (''Acanthocybium solandri'') is a scombrid fish found worldwide in tropical and subtropical seas. In Hawaii, the wahoo is known as ono. The species is sometimes called hoo in the United States. It is best known to sports fishermen, a ...

(''Acanthocybium solandri'') has been known to reach speeds of up to .

Distribution

Most mackerel species have restricted distribution ranges, while others, like the Wahoo, are found in all oceans between 59°N and 48°S. In combination, these fish (Scombroids) can be found worldwide in all oceans of the world, excluding the Southern and Arctic Oceans. However, the Atlantic chub mackerel has been spotted near

Greenland Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...

in the

Arctic Ocean The Arctic Ocean is the smallest and shallowest of the world's five oceanic divisions. It spans an area of approximately and is the coldest of the world's oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, ...

on rare occasions as

global warming Present-day climate change includes both global warming—the ongoing increase in global average temperature—and its wider effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes ...

makes the Arctic more habitable for them. Some mackerel species, especially

pelagic The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...

and

filter feeding Filter feeders are aquatic animals that acquire nutrients by feeding on organic matters, food particles or smaller organisms (bacteria, microalgae and zooplanktons) suspended in water, typically by having the water pass over or through a spe ...

one such as chub mackerel, migrate vertically, staying at certain depths depending on the time of day and food availability. For example, adult snake mackerel, conduct a diel vertical migration, staying in deeper water during the day and rising to the surface at night to feed. The young and juveniles also migrate vertically, but in the opposite direction, staying near the surface during the day and moving deeper at night.

Lifecycle and ecology (Scombroids)

Mackerel are prolific broadcast spawners, and must breed near the surface of the water because the eggs of the females float. Individual females lay between 300,000 and 1,500,000 eggs. Their eggs and

larvae A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect developmental biology, development such as insects, some arachnids, amphibians, or cnidarians typical ...

are

, that is, they float free in the open sea. The larvae and juvenile mackerel feed on

zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...

. As adults, they have sharp teeth, and hunt small crustaceans such as

copepod Copepods (; meaning 'oar-feet') are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (living in the water column), some are benthos, benthic (living on the sedimen ...

shrimp A shrimp (: shrimp (American English, US) or shrimps (British English, UK)) is a crustacean with an elongated body and a primarily Aquatic locomotion, swimming mode of locomotion – typically Decapods belonging to the Caridea or Dendrobranchi ...

, and

squid A squid (: squid) is a mollusc with an elongated soft body, large eyes, eight cephalopod limb, arms, and two tentacles in the orders Myopsida, Oegopsida, and Bathyteuthida (though many other molluscs within the broader Neocoleoidea are also ...

. In turn, they are hunted by larger pelagic animals such as tuna,

billfish The billfish are a group (Xiphioidea) of saltwater fish, saltwater predatory fish characterised by prominent pointed beak, bills (rostrum (anatomy), rostra), and by their large size; some are longer than . Extant billfish include sailfish and m ...

, sea lions,

shark Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the ...

s, and

pelican Pelicans (genus ''Pelecanus'') are a genus of large water birds that make up the family Pelecanidae. They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before ...

s. Off

Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...

, spinner sharks follow migrating schools of mackerel.

Bryde's whale Bryde's whale ( ), or the Bryde's whale complex, putatively comprises three species of rorqual and possibly four. The "complex" means the number and classification remain unclear because of a lack of definitive information and research. The c ...

s feed on mackerel when they can find them. They use several feeding methods, including skimming the surface, lunging, and

bubble net Bubble-net feeding is a feeding behavior engaged in by humpback whale, humpback whales and Bryde's whale, Bryde's whales. It is one of the few Cetacean surfacing behaviour, surface feeding behaviors that humpback whales are known to engage in. Th ...

Fisheries

A bunch of Mackerel freshly caught on a African beach

Chub mackerel, ''Scomber japonicus'', are the most intensively fished scombroid mackerel. They account for about half the total capture production of scombroid mackerels. As a species, they are easily confused with Atlantic mackerel. Chub mackerel migrate long distances in oceans and across the Mediterranean. They can be caught with drift nets and suitable trawls, but are most usually caught with surround nets at night by attracting them with lampara lamps. The remaining catch of scombroid mackerels is divided equally between the Atlantic mackerel and all other scombroid mackerels. Just these two species (Chub mackerel and Atlantic mackerel) account for about 75% of the total catch of scombroid mackerels. Chilean jack mackerel are the most commonly fished non-scombroid mackerel, fished as heavily as chub mackerel. The species has been

overfished Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in the ...

, and its fishery may now be in danger of collapsing. Smaller mackerel behave like

herring Herring are various species of forage fish, belonging to the Order (biology), order Clupeiformes. Herring often move in large Shoaling and schooling, schools around fishing banks and near the coast, found particularly in shallow, temperate wate ...

s, and are captured in similar ways. Fish species like these, which

school A school is the educational institution (and, in the case of in-person learning, the Educational architecture, building) designed to provide learning environments for the teaching of students, usually under the direction of teachers. Most co ...

near the surface, can be caught efficiently by purse seining. Huge purse-seine vessels use spotter planes to locate the schooling fish. Then they close in using sophisticated sonar to track the shape of the school, which is then encircled with fast auxiliary boats that deploy purse seines as they speed around the school. Suitably designed trollers can also catch mackerels effectively when they swim near the surface. Trollers typically have several long booms which they lift and drop with "topping lifts". They haul their lines with electric or hydraulic reels.

Fish aggregating device A fish aggregating (or aggregation) device (FAD) is a man-made object used to attract pelagic fish such as marlin, tuna and mahi-mahi (dolphin fish). They usually consist of buoys or floats tethered to the ocean floor. Various types of FADs have be ...

s are also used to target mackerel.

Management and conservation

The

North Sea The North Sea lies between Great Britain, Denmark, Norway, Germany, the Netherlands, Belgium, and France. A sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian Se ...

has been overfished to the point where the ecological balance has become disrupted and many jobs in the fishing industry have been lost. The Southeast US region spans the

Gulf of Mexico The Gulf of Mexico () is an oceanic basin and a marginal sea of the Atlantic Ocean, mostly surrounded by the North American continent. It is bounded on the northeast, north, and northwest by the Gulf Coast of the United States; on the southw ...

, the

Caribbean Sea The Caribbean Sea is a sea of the Atlantic Ocean, North Atlantic Ocean in the tropics of the Western Hemisphere, located south of the Gulf of Mexico and southwest of the Sargasso Sea. It is bounded by the Greater Antilles to the north from Cuba ...

, and the US Southeast Atlantic. Overfishing of

king King is a royal title given to a male monarch. A king is an Absolute monarchy, absolute monarch if he holds unrestricted Government, governmental power or exercises full sovereignty over a nation. Conversely, he is a Constitutional monarchy, ...

and

occurred in the 1980s. Regulations were introduced to restrict the size, fishing locations, and bag limits for recreational fishers and commercial fishers.

Gillnet Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...

s were banned in waters off Florida. By 2001, the mackerel stocks had bounced back.

As food

Mackerel is an important

food fish Many species of fish are caught by humans and consumed as food in virtually all regions around the world. Their meat has been an important dietary source of protein and other nutrients in the human diet. The English language does not have a s ...

that is consumed worldwide. As an

oily fish Oily fish are fish species with fish oil, oil (fats) in soft tissues and in the coelomic cavity around the Gut (zoology), gut. Their fillet (cut), fillets may contain up to 30% oil, although this figure varies both within and between species. ...

, it is a rich source of

omega-3 fatty acid Omega−3 fatty acids, also called omega−3 oils, ω−3 fatty acids or ''n''−3 fatty acids, are polyunsaturated fatty acids (PUFAs) characterized by the presence of a double bond three atoms away from the terminal methyl group in their ...

s. The flesh of mackerel spoils quickly, especially in the tropics, and can cause scombroid food poisoning. Accordingly, it should be eaten on the day of capture, unless properly refrigerated or cured. Mackerel preservation is not simple. Before the 19th-century development of canning and the widespread availability of refrigeration, salting and smoking were the principal preservation methods available. Historically in England, this fish was not preserved, but was consumed only in its fresh form. However, spoilage was common, leading the authors of ''The Cambridge Economic History of Europe'' to remark: "There are more references to stinking mackerel in English literature than to any other fish!" In France, mackerel was traditionally pickled with large amounts of salt, which allowed it to be sold widely across the country. For many years mackerel was regarded as 'unclean' in the UK and other places due to folklore which suggested that the fish fed on the corpses of dead sailors. A 1976 survey of housewives in Britain undertaken by the White Fish Authority indicated a reluctance to departing from buying the traditional staples of cod, haddock or salmon. Less than 10% of the survey's 1,931 respondents had ever bought mackerel, and only 3% did so regularly. As a result of this trend, many UK

fishmonger A fishmonger (historically fishwife for female practitioners) is someone who sells raw fish and seafood. Fishmongers can be wholesalers or retailers and are trained at selecting and purchasing, handling, gutting, boning, filleting, displaying, ...

s during the 1970s did not display or even stock mackerel.

References

External links

''British Marine Life Study Society''. Retrieved 3 March 2012.
Mackerel nutrition facts

* {{commercial fish topics Scombridae Oily fish Commercial fish Fish common names