The southern pig-tailed macaque (''Macaca nemestrina''), also known as the Sundaland pig-tailed macaque and the Sunda pig-tailed macaque, is a medium-sized

macaque The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and Europe (in Gibraltar). Macaques are principally f ...

that lives in

Sundaland Sundaland (also called Sundaica or the Sundaic region) is a biogeographical region of Southeast Asia corresponding to a larger landmass that was exposed throughout the last 2.6 million years during periods when sea levels were lower. It inc ...

southern Thailand Southern Thailand (formerly Southern Siam and Tambralinga) is the southernmost cultural region of Thailand, separated from Central Thailand by the Kra Isthmus. Geography Southern Thailand is on the Malay Peninsula, with an area of around , bo ...

Malaysia Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...

, and

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...

. It is known locally as beruk.

Etymology and taxonomy

The

species epithet Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In Taxonomy (biology), taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany) ...

, ''nemestrina'', is an adjective (derived from Latin , meaning "the god of groves") modified to agree in gender with the feminine generic name. ''M. nemestrina'' formerly included the northern pig-tailed, Pagai Island, and Siberut macaques as

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

. All four are now considered separate species. In the 19th century, ''bruh'' was the native name used by Malays in Sumatra for the macaque.

Description

As with other ''Macaca'' species, males are larger than females; while males are measured at in length and in weight, females are measured at in length and in weight. This macaque has buff-brown fur, with a darker dorsal area and lighter ventral area. Its common name refers to the short tail held semi-erect, resembling the tail of a pig.

Behaviour and ecology

''M. nemestrina'' is mainly

terrestrial Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...

, but also a skilled climber. Unlike almost all primates, these macaques love water. They live in large groups that split into smaller groups during

daytime Daytime or day as observed on Earth is the period of the day during which a given location experiences Daylight, natural illumination from direct sunlight. Daytime occurs when the Sun appears above the local horizon, that is, anywhere on the ...

when they are

foraging Foraging is searching for wild food resources. It affects an animal's fitness because it plays an important role in an animal's ability to survive and reproduce. Foraging theory is a branch of behavioral ecology that studies the foraging behavi ...

. They are

omnivorous An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize ...

, feeding mainly on fruits, seeds, berries,

cereal A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...

fungi A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...

, and

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

s. A study in peninsular

found them to be the primary, and perhaps the only, seed dispersers of the

rattan Rattan, also spelled ratan (from Malay language, Malay: ''rotan''), is the name for roughly 600 species of Old World climbing palms belonging to subfamily Calamoideae. The greatest diversity of rattan palm species and genera are in the clos ...

species ''

Calamus calicarpus Calamus may refer to: Botany and zoology * ''Calamus'' (fish), a genus of fish in the family Sparidae * ''Calamus'' (palm), a genus of rattan palms * Calamus, the hollow shaft of a feather, also known as the quill * ''Acorus calamus'', the swee ...

'' (syn. ''Daemonorops calicarpa'') and ''

Calamus castaneus Calamus may refer to: Botany and zoology * ''Calamus'' (fish), a genus of fish in the family Sparidae * ''Calamus'' (palm), a genus of rattan palms * Calamus, the hollow shaft of a feather, also known as the quill * ''Acorus calamus'', the swee ...

''. There is a

hierarchy A hierarchy (from Ancient Greek, Greek: , from , 'president of sacred rites') is an arrangement of items (objects, names, values, categories, etc.) that are represented as being "above", "below", or "at the same level as" one another. Hierarchy ...

among males, based on strength, and among females, based on

heredity Heredity, also called inheritance or biological inheritance, is the passing on of traits from parents to their offspring; either through asexual reproduction or sexual reproduction, the offspring cells or organisms acquire the genetic infor ...

. Thus, the daughter of the

alpha Alpha (uppercase , lowercase ) is the first letter of the Greek alphabet. In the system of Greek numerals, it has a value of one. Alpha is derived from the Phoenician letter ''aleph'' , whose name comes from the West Semitic word for ' ...

female will immediately be placed above all other females in the group. The alpha female leads the group, while the male role is more to manage conflict within the group and to defend it. Female

gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...

lasts around 5.7 months. She will give birth to one infant every two years. Weaning occurs at 4–5 months.

Sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...

is reached at 3–5 years. In

Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...

, they have been trained for 400 years to harvest

coconut The coconut tree (''Cocos nucifera'') is a member of the palm tree family (biology), family (Arecaceae) and the only living species of the genus ''Cocos''. The term "coconut" (or the archaic "cocoanut") can refer to the whole coconut palm, ...

Habitat and distribution

This

is mostly found in rainforest up to , but will also enter

plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...

s and gardens. It is found in the southern half of the

Malay Peninsula The Malay Peninsula is located in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area contains Peninsular Malaysia, Southern Tha ...

(only just extending into southernmost

Borneo Borneo () is the List of islands by area, third-largest island in the world, with an area of , and population of 23,053,723 (2020 national censuses). Situated at the geographic centre of Maritime Southeast Asia, it is one of the Greater Sunda ...

Sumatra Sumatra () is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the list of islands by area, sixth-largest island in the world at 482,286.55 km2 (182,812 mi. ...

and

Bangka Island Bangka is an island lying east of Sumatra, Indonesia. It is administered under the province of the Bangka Belitung Islands, being one of its namesakes alongside the smaller island of Belitung across the Gaspar Strait. The 9th largest island in ...

. There are reports of the species having been present in

Singapore Singapore, officially the Republic of Singapore, is an island country and city-state in Southeast Asia. The country's territory comprises one main island, 63 satellite islands and islets, and one outlying islet. It is about one degree ...

before 1950, but these were likely escaped pets. The only pig-tailed macaques in Singapore today are introduced monkeys.

Interaction with humans

Since the 19th century, monkeys including the southern pig-tailed macaque, have been used by humans to harvest coconuts in southeast Asia.

References

External links

* ARKive
images and movies of the Sunda pig-tailed macaque ''(Macaca nemestrina)''Biolib
{{Taxonbar, from=Q618026 Macaca Primates of Southeast Asia Mammals described in 1766 Taxa named by Carl Linnaeus Fauna of the Borneo lowland rain forests Primates of Borneo