Lysimachia
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''Lysimachia'' ( ) is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
consisting of 182 accepted species of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s traditionally classified in the family
Primulaceae The Primulaceae ( ), commonly known as the primrose family (but not related to the Onagraceae, evening primrose family), are a family (biology), family of Herbaceous plant, herbaceous and woody flowering plants including some favourite garden pla ...
. Based on a
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study it was transferred to the family Myrsinaceae, before this family was later merged into the Primulaceae.


Characteristics

''Lysimachia'' species often have yellow flowers, and grow vigorously. They tend to grow in damp conditions. Several species within ''Lysimachia'' are commonly called loosestrife, although this name is also used for plants within the genus '' Lythrum''. The genus is named in honor of
Lysimachus Lysimachus (; Greek language, Greek: Λυσίμαχος, ''Lysimachos''; c. 360 BC – 281 BC) was a Thessaly, Thessalian officer and Diadochi, successor of Alexander the Great, who in 306 BC, became king of Thrace, Anatolia, Asia Minor and Mace ...
, a king of ancient Sicily, who is said to have calmed a mad ox by feeding it a member of the genus. ''Lysimachia'' species are used as food plants by the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
e of some butterflies and moths, including the dot moth, grey pug, lime-speck pug,
small angle shades The small angle shades (''Euplexia lucipara'') is a moth of the family Noctuidae. It is distributed throughout the Palearctic. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. As the common name ...
, and v-pug.


Specialized pollinators

Bees of the genus '' Macropis'' are specialized to pollinate oil-producing ''Lysimachia'' plants. These bees use exclusively ''Lysimachia'' floral oils for building their nests and provisioning cells. ''Lysimachia'' floral-specific chemicals are strong attractors for '' Macropis nuda'' and '' Macropis fulvipes'' bees that are seldom found in other plant genera.


Fossil record

Several
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
seeds of ''Lysimachia '' sp. have been described from
middle Miocene The Middle Miocene is a sub-epoch of the Miocene epoch (geology), epoch made up of two Stage (stratigraphy), stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0. ...
strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...
of the Fasterholt area near
Silkeborg Silkeborg () is a Denmark, Danish town with a population of 52,571 (1 January 2025).Jutland Jutland (; , ''Jyske Halvø'' or ''Cimbriske Halvø''; , ''Kimbrische Halbinsel'' or ''Jütische Halbinsel'') is a peninsula of Northern Europe that forms the continental portion of Denmark and part of northern Germany (Schleswig-Holstein). It ...
,
Denmark Denmark is a Nordic countries, Nordic country in Northern Europe. It is the metropole and most populous constituent of the Kingdom of Denmark,, . also known as the Danish Realm, a constitutionally unitary state that includes the Autonomous a ...
. †''Lysimachia nikitinii'' seed
fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
have been collected from
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...
of south eastern
Belarus Belarus, officially the Republic of Belarus, is a landlocked country in Eastern Europe. It is bordered by Russia to the east and northeast, Ukraine to the south, Poland to the west, and Lithuania and Latvia to the northwest. Belarus spans an a ...
. The fossils are most similar to seeds of the
East Asian East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...
'' Lysimachia davurica''.The
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Belarus Belarus, officially the Republic of Belarus, is a landlocked country in Eastern Europe. It is bordered by Russia to the east and northeast, Ukraine to the south, Poland to the west, and Lithuania and Latvia to the northwest. Belarus spans an a ...
and its correlation with other
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
by Felix Yu. VELICHKEVICH and Ewa ZASTAWNIAK - Acta Palaeobotanica 43(2): 137–259, 2003


Selected species

282 species are accepted. Selected species include: *''
Lysimachia arvensis ''Lysimachia arvensis'', synonym (taxonomy), syn. ''Anagallis arvensis'', commonly known as scarlet pimpernel, red pimpernel, red chickweed, poor man's barometer, poor man's weather-glass, shepherd's weather glass or shepherd's clock, is a spec ...
'' L. – Scarlet pimpernel *'' Lysimachia asperulifolia'' , orth. var. ''L. asperulaefolia'' – roughleaf yellow loosestrife (endemic to Atlantic coastal plain in North and South Carolina) *'' Lysimachia atropurpurea'' L. – purple gooseneck loosestrife *'' Lysimachia azorica'' Hornem. ex Hook. – (endemic to the Azores) *'' Lysimachia barystachys'' Bunge – Manchurian yellow loosestrife *'' Lysimachia ciliata'' L. – fringed loosestrife (North America) *'' Lysimachia clethroides'' Duby – gooseneck loosestrife * ''Lysimachia'' × ''commixta'' – a hybrid of ''L. terrestris'' and ''L. thyrsiflora'' * '' Lysimachia congestiflora'' *'' Lysimachia daphnoides'' (A.Gray) Hillebr. – ''Lehua makanoe'' (Island of
Kauai Kauai (), anglicized as Kauai ( or ), is one of the main Hawaiian Islands. It has an area of 562.3 square miles (1,456.4 km2), making it the fourth-largest of the islands and the 21st-largest island in the United States. Kauai lies 73 m ...
in
Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...
) *'' Lysimachia filifolia'' C.N.Forbes (Islands of
Oahu Oahu (, , sometimes written Oahu) is the third-largest and most populated island of the Hawaiian Islands and of the U.S. state of Hawaii. The state capital, Honolulu, is on Oahu's southeast coast. The island of Oahu and the uninhabited Northwe ...
and Kauai in Hawaii) *'' Lysimachia foenum-graecum'' Hance *'' Lysimachia fraseri'' Duby – Fraser's yellow loosestrife (Southeastern United States) *'' Lysimachia glutinosa'' *'' Lysimachia hillebrandii'' Hook.f. ex A.Gray – ''kolokolo kuahiwi'' (Hawaii) *'' Lysimachia hybrida'' Michx. *'' Lysimachia iniki'' – Wailua River yellow loosestrife (endemic to Kauai, Hawaii) *'' Lysimachia japonica'' *'' Lysimachia latifolia'' – Pacific starflower *'' Lysimachia lichiangensis'' Forrest *'' Lysimachia lydgatei'' – Maui yellow loosestrife (endemic to Maui, Hawaii) *'' Lysimachia maritima'' *'' Lysimachia mauritiana'' *'' Lysimachia minoricensis'' J.J.Rodr. (
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
) *'' Lysimachia nemorum'' L. – yellow pimpernel *'' Lysimachia nummularia'' L. – creeping jenny, moneywort (Europe, introduced in North America) *'' Lysimachia pendens'' * ''Lysimachia'' × ''producta'' – a hybrid of ''L. terrestris'' and ''L. quadrifolia'' *'' Lysimachia punctata'' L. – spotted loosestrife *'' Lysimachia quadriflora'' Sims – four-flower yellow loosestrife (Eastern North America) *'' Lysimachia quadrifolia'' L. – whorled loosestrife (Eastern North America) *'' Lysimachia remyi'' (islands of
Maui Maui (; Hawaiian language, Hawaiian: ) is the second largest island in the Hawaiian archipelago, at 727.2 square miles (1,883 km2). It is the List of islands of the United States by area, 17th-largest in the United States. Maui is one of ...
and
Molokai Molokai or Molokai ( or ; Molokaʻi dialect: Morotaʻi ) is the fifth most populated of the eight major islands that make up the Hawaiian Islands archipelago in the middle of the Pacific Ocean. It is 38 by 10 miles (61 by 16 km) at its g ...
in Hawaii) ** ''Lysimachia remyi'' subsp. ''maxima'' (Masam.) I.C.Oh & Anderb. (Molokai) **''Lysimachia remyi'' subsp. ''remyi'' (Maui and Molokai) *'' Lysimachia scopulensis'' *'' Lysimachia sertulata'' Baudo – Chilean melilukul *'' Lysimachia terrestris'' (L.) Britton, Sterns & Poggenb. – swamp candles (North America) *'' Lysimachia thyrsiflora'' – tufted loosestrife *'' Lysimachia venosa'' – veined yellow loosestrife (endemic to Kauai, Hawaii) *'' Lysimachia verticillaris'' Biehler *'' Lysimachia vulgaris'' L. – garden loosestrife, yellow loosestrife (Eurasia)


References


External links


Lysimachus' Dog & Nisaean Horses
- Informative but non-scholarly essay on Lysimachia & Lysimachus (Annotated with Sources). *Pictures o


PAF - Lysimachia clethroides - Duby.
{{Authority control Primulaceae genera Taxa named by Carl Linnaeus