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''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family
Lamiaceae The Lamiaceae ( ) or Labiatae are a family of flowering plants commonly known as the mint, deadnettle or sage family. Many of the plants are aromatic in all parts and include widely used culinary herbs like basil, mint, rosemary, sage, savo ...
. They are all
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition ...
plants native to
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
,
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
, Australia, and North America.Kew World Checklist of Selected Plant Families
/ref> The species are most often found in wetlands, damp meadows, and stream banks. Some of the wetland species have become endangered.


Appearance

The genus includes only
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
species; they spread by both
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s and stems rooting as they grow along the ground. Small white flowers bloom in late summer on leaf axials. Leaves are bright green, pointed, lobed, and like all mints occur in opposite pairs. Some species start with curled purple leaves that unfurl to a bright green coloration. The species in this genus vary in size, but generally grow to about .


Ethnobotanical history

The plant's juice yields black
dye A dye is a colored substance that chemically bonds to the substrate to which it is being applied. This distinguishes dyes from pigments which do not chemically bind to the material they color. Dye is generally applied in an aqueous solution and ...
, supposedly used by the
Roma Roma or ROMA may refer to: Places Australia * Roma, Queensland, a town ** Roma Airport ** Roma Courthouse ** Electoral district of Roma, defunct ** Town of Roma, defunct town, now part of the Maranoa Regional Council * Roma Street, Brisbane, a ...
in
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
to tan their skin to mimic Egyptians, hence the common name of ''Gypsywort'' for '' Lycopus europaeus.''
Apothecaries ''Apothecary'' () is a mostly archaic term for a medical professional who formulates and dispenses '' materia medica'' (medicine) to physicians, surgeons, and patients. The modern chemist (British English) or pharmacist (British and North Amer ...
and herbalists used the leaves, stems, and flowers for their
astringent An astringent (sometimes called adstringent) is a chemical that shrinks or constricts body tissues. The word derives from the Latin ''adstringere'', which means "to bind fast". Calamine lotion, witch hazel, and yerba mansa, a Californian pl ...
and sedative qualities as well as for anxiety,
tuberculosis Tuberculosis (TB) is an infectious disease usually caused by ''Mycobacterium tuberculosis'' (MTB) bacteria. Tuberculosis generally affects the lungs, but it can also affect other parts of the body. Most infections show no symptoms, in w ...
, and palpitations.


Fossil record

Fossil seeds of †''Lycopus antiquus'' are known from the Middle
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
strata In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as e ...
of southern
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
, from the Miocene of
Lower Lusatia Lower Lusatia (; ; ; szl, Dolnŏ Łużyca; ; ) is a historical region in Central Europe, stretching from the southeast of the German state of Brandenburg to the southwest of Lubusz Voivodeship in Poland. Like adjacent Upper Lusatia in the ...
,
Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
, and from the Late Miocene
strata In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as e ...
of western
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...
and
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian invas ...
. ''Lycopus antiquus'' has possibly been applied to more than one extinct species which were widely distributed in Europe and
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...
from the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
to the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Lycopus alissoviae ''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family Lamiaceae. They are all herbaceous plants native to Europe, Asia, Australia, and North America. The species are most often found in wetlands, damp meadows, and str ...
'' Prob. – Primorye region of Russia #''
Lycopus americanus ''Lycopus americanus'', common names American water horehound or American bugleweed, is a member of the genus '' Lycopus''. It blooms in late summer and is found in much of North America. Medicinal plant It is reputed to have medicinal properti ...
'' Muhl. ex W.P.C.Barton – American bugleweed - widespread across most of United States and Canada #''
Lycopus amplectens Lycopus amplectens, common names clasping-leaved water-horehound, sessile-leaved bugleweed, and sessile-leaved water-horehound, is a species of '' Lycopus'' native to North America. Conservation status in the United States It is listed as endan ...
'' Raf. – eastern United States #''
Lycopus angustifolius ''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family Lamiaceae. They are all herbaceous plants native to Europe, Asia, Australia, and North America. The species are most often found in wetlands, damp meadows, and str ...
'' Elliott – southeastern United States #''
Lycopus asper ''Lycopus asper'' is a species of flowering plant in the mint family known by the common name rough bugleweed. It is native to much of North America, where it can be found most often in moist areas, such as the soil near lakes. This is a perenni ...
'' Greene – rough bugleweed - western Canada, western + central United States #'' Lycopus australis'' R.Br. – Australian Gypsywort - Australia #''
Lycopus cavaleriei ''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family Lamiaceae. They are all herbaceous plants native to Europe, Asia, Australia, and North America. The species are most often found in wetlands, damp meadows, and strea ...
'' H.Lév. – Korean bugleweed - China, Japan, Korea, Sakhalin, Kuril Islands #''
Lycopus charkeviczii ''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family Lamiaceae. They are all herbaceous plants native to Europe, Asia, Australia, and North America. The species are most often found in wetlands, damp meadows, and str ...
'' Prob. – Primorye region of Russia #''
Lycopus cokeri ''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family Lamiaceae. They are all herbaceous plants native to Europe, Asia, Australia, and North America. The species are most often found in wetlands, damp meadows, and str ...
'' H.E.Ahles ex Sorrie – North Carolina, South Carolina #'' Lycopus europaeus'' L. – Gypsywort - Europe, North Africa, northern Asia; naturalized in New Zealand and North America #'' Lycopus exaltatus'' L.f. – central + eastern Europe, Siberia, Central Asia, Xinjiang, Caucasus, Western Himalayas #''
Lycopus hirtellus ''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family Lamiaceae. They are all herbaceous plants native to Europe, Asia, Australia, and North America. The species are most often found in wetlands, damp meadows, and str ...
'' Kom. – Primorye region of Russia #''
Lycopus × intermedius ''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family Lamiaceae. They are all herbaceous plants native to Europe, Asia, Australia, and North America. The species are most often found in wetlands, damp meadows, and strea ...
'' Hausskn. – Germany, Austria, Czech Republic, Greece ''(L. europaeus × L. exaltatus)'' #''
Lycopus kurilensis ''Lycopus'' (water horehound, gypsywort, or bugleweed) is a genus in the family Lamiaceae. They are all herbaceous plants native to Europe, Asia, Australia, and North America. The species are most often found in wetlands, damp meadows, and str ...
'' Prob. – Kuril Islands #'' Lycopus laurentianus'' Roll.-Germ. – Quebec #''
Lycopus lucidus ''Lycopus lucidus'' is a species of plant in the family Lamiaceae. The aerial parts (澤蘭, 泽兰 zélán) and the dried rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and sh ...
'' Turcz. ex Benth.
Traditional Chinese Medicine Traditional Chinese medicine (TCM) is an alternative medicine, alternative medical practice drawn from traditional medicine in China. It has been described as "fraught with pseudoscience", with the majority of its treatments having no logica ...
herb to make ''
Lycopi rhizoma ''Lycopus lucidus'' is a species of plant in the family Lamiaceae. The aerial parts (澤蘭, 泽兰 zélán) and the dried rhizomes (地筍, 地笋 dìsǔn) are used in traditional Chinese medicine Traditional Chinese medicine (TCM) is a ...
'' - China, Japan, Korea, Siberia, Russian Far East #'' Lycopus rubellus'' Moench – central + eastern United States #'' Lycopus × sherardii'' Steele – Quebec, Ontario, eastern United States #'' Lycopus sichotensis'' Prob. – Primorye region of Russia #''
Lycopus uniflorus ''Lycopus uniflorus'' is a species of flowering plant in the mint family known by the common name northern bugleweed. It is native to much of North America (Canada, United States) and east Asia (China, Japan, Korea, Russian Far East) ''Lycopus ...
'' Michx. – northern bugleweed - Canada, United States. China, Japan, Korea, Russian Far East #'' Lycopus virginicus'' L. – Virginia bugleweed/water-horehound - central + eastern United States


References

{{Taxonbar, from=Q158341 Lamiaceae genera Taxa named by Carl Linnaeus