''Lycophocyon'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

caniformia Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include Canidae, dogs (Wolf, wolves, foxes, etc.), bears, raccoons, and Mustelidae, mustelids. The Pinnipedia (pinniped, seals, walruses and sea lions) ...

carnivora Carnivora ( ) is an order of placental mammals specialized primarily in eating flesh, whose members are formally referred to as carnivorans. The order Carnivora is the sixth largest order of mammals, comprising at least 279 species. Carnivor ...

n from

Middle Eocene The Eocene ( ) is a geological epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''Ēṓs'', ' Dawn') a ...

(early Duchesnean and possibly late

Uintan The Uintan North American Stage is the North American faunal stage, typically set from 46,200,000 to 42,000,000 years before present lasting 4.2 million years. The Uintan Stage is a key part of the North American land mammal age, North American Lan ...

NALMA) deposits of

San Diego County San Diego County (), officially the County of San Diego, is a county in the southwest corner of the U.S. state of California, north to its border with Mexico. As of the 2020 census, the population was 3,298,634; it is the second-most populous ...

California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...

. ''Lycophocyon'' is known from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

UCMP 85202, a partial left and right

dentary In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone ...

. Paratypes include UCMP 170713, SDSNH 107658, SDSNH 107659, SDSNH 107442, SDSNH 107443 and SDSNH 107444, partial dentaries,

mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...

s and other

cranial Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek language, Greek roots, describe something in its standard anatomical position. Thi ...

remains, and SDSNH 107446 and SDSNH 107447, cranial and

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

l fragments. Many additional specimens are also known. All specimens were collected from numerous localities, all of them from the upper portions of "member C" of the Santiago Formation. It was first named by Susumu Tomiya in

2011 The year marked the start of a Arab Spring, series of protests and revolutions throughout the Arab world advocating for democracy, reform, and economic recovery, later leading to the depositions of world leaders in Tunisia, Egypt, and Yemen ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Lycophocyon hutchisoni''. The generic name means "twilight dog" in

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

, in references to its occurrence on the west coast of North America, and its affinity to the

Caniformia Caniformia is a suborder within the order Carnivora consisting of "dog-like" carnivorans. They include Canidae, dogs (Wolf, wolves, foxes, etc.), bears, raccoons, and Mustelidae, mustelids. The Pinnipedia (pinniped, seals, walruses and sea lions) ...

. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

honours the paleontologist J. Howard Hutchison.

Phylogeny

The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below follows the topology from a 2011 analysis by Susumu Tomiya.

References

Eocene carnivorans Fossil taxa described in 2011 Extinct mammals of North America Prehistoric carnivoran genera {{paleo-carnivora-stub