Lycalopex Gymnocercus
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The Pampas fox (''Lycalopex gymnocercus''), also known as grey pampean fox, Pampas zorro, Azara's fox, or Azara's zorro (in Guaraní also called , anglicized as aguarachay, in Portuguese also called (), is a medium-sized
zorro Zorro ( or , Spanish for "fox") is a fictional character created in 1919 by American Pulp magazine, pulp writer Johnston McCulley, appearing in works set in the Pueblo de Los Ángeles in Alta California. He is typically portrayed as a dashin ...
, or "false" fox, native to the South American
Pampas The Pampas (; from Quechua 'plain'), also known as the Pampas Plain, are fertile South American low grasslands that cover more than and include the Argentine provinces of Buenos Aires, La Pampa, Santa Fe, Entre Ríos, and Córdoba; all o ...
. ''Azara'' in some of its alternative common names is a reference to
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many countries in the Americas **Spanish cuisine **Spanish history **Spanish culture ...
naturalist Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
Félix de Azara Félix de Azara y Perera (18 May 1742 – 20 October 1821) was a Spanish military officer, naturalist, and engineer. Life Félix de Azara y Perera was born on 18 May 1746 in Barbuñales, Aragon. He joined the army and attended a Spanish milita ...
.


Description

The Pampas fox resembles the
culpeo The culpeo (''Lycalopex culpaeus''), also known as Culpeo zorro, Andean zorro, Andean fox, Paramo wolf, Andean wolf,Comparative ecology of two South American foxes, 'Dusicvon ariseus' and 'Culpaeus' by Warren E. Johnson. Doctoral dissertation. Io ...
or Andean fox in appearance and size, but has a proportionately wider snout, reddish fur on the head and neck, and a black mark on the muzzle. Its short, dense fur is grey over most of the body, with a black line running down the back and onto the tail, and pale, almost white, underparts. The ears are triangular, broad, and relatively large, and are reddish on the outer surface and white on the inner surface. The inner surfaces of the legs are similar in color to the underparts, while the outer surface is reddish on the fore limbs, and grey on the hind limbs; the lower hind limb also bears a distinctive black spot. Adults range from in body length, and weigh ; males are about 10% heavier than females. In the northern part of its range, the pampas fox is more richly colored than in the southern part.


Distribution and habitat

The Pampas fox can be found primarily in northern and central
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
,
Uruguay Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
, eastern
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
,
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
, and southern
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
. It prefers open pampas habitats, often close to agricultural land, but can also be found in montane or chaco forest, dry scrubland, and wetland habitats. It is most common below elevation, but can inhabit puna grasslands up to . Five subspecies of ''L. gymnocercus'' are generally recognized *''L. g. antiquus'' (Ameghino, 1889:298); — Found in the Pampas grasslands, Monte shrublands and Espinal open woodlands of central Argentina, from Córdoba and San Luis Provinces to the Río Negro and the Atlantic coast. *''L. g. domeykoanus'' (Philippi, 1901:168); Found in Copiapó Province, Chile. *''L. g. gracilis'' (Burmeister, 1861:406); Found in the Pampas surrounding Mendoza, Argentina. *''L. g. gymnocercus'' (Fischer, 1814:178); Found in the subtropical grasslands of northeastern Argentina, Uruguay, Paraguay, Bolivia and eastern Brazil. *''L. g. maullinicus'' (Philippi, 1903:158); Found in Llanquihue Province, Chile, east of Llanquihue Lake An earlier (1982) taxonomical revision recognized only and , along with the otherwise unaccepted subspecies (Massoia, 1982:149), which is restricted to the Chaco-Yungas Mountain Tropical Forest in Salta and Jujuy Provinces. That revision classified , , and as subspecies of , in part due to their falling outside the known present range of . Fossils of this species are known from the late
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
in Argentina.


Local names

In the Spanish-speaking areas of its habitat, the Pampas fox is known by the common names of ''zorro de las pampas'' or ''zorro gris pampeano''. In Portuguese-speaking Brazil, it is called by the common names of ''graxaim'' or ''sorro''.


Behavior and diet

The Pampas foxes mostly live a solitary life, but come together as
monogamous Monogamy ( ) is a relationship of two individuals in which they form a mutual and exclusive intimate partnership. Having only one partner at any one time, whether for life or serial monogamy, contrasts with various forms of non-monogamy (e.g. ...
pairs in the
breeding season Seasonal breeders are animal species that successfully mate only during certain times of the year. These times of year allow for the optimization of survival of young due to factors such as ambient temperature, food and water availability, and ch ...
to raise their young. They are mainly
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
, becoming active at dusk, although may also be active during the day. They den in any available cavity, including caves, hollow trees, and the burrows of
viscacha Viscacha or vizcacha (, ) are rodents of two genera ('' Lagidium'' and '' Lagostomus'') in the family Chinchillidae. They are native to South America and convergently resemble rabbits. The five extant species of viscacha are: *The Plains vi ...
s or
armadillo Armadillos () are New World placental mammals in the order (biology), order Cingulata. They form part of the superorder Xenarthra, along with the anteaters and sloths. 21 extant species of armadillo have been described, some of which are dis ...
s. Even when raising young together, adult foxes generally hunt alone, marking their territory by defecating at specific latrine sites. Although considerable variation is seen, the home range of a typical Pampas fox has been estimated to be around . Pampas foxes are more
omnivorous An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize ...
than most other canids, and have a varied and opportunistic diet. Their primary prey consists of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s,
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s,
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores and live Solitary animal, solitarily or in pairs. They nest in slight depressions called forms, and their young are precociality, able to fend for themselves ...
s,
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
,
carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
, and
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s, although they also eat
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s,
armadillo Armadillos () are New World placental mammals in the order (biology), order Cingulata. They form part of the superorder Xenarthra, along with the anteaters and sloths. 21 extant species of armadillo have been described, some of which are dis ...
s,
snail A snail is a shelled gastropod. The name is most often applied to land snails, terrestrial molluscs, terrestrial pulmonate gastropod molluscs. However, the common name ''snail'' is also used for most of the members of the molluscan class Gas ...
s and other invertebrates, lambs, and the eggs of ground-nesting birds. Their primary predators are pumas, domesticated dogs, and anacondas, most notably the yellow anaconda.


Reproduction

Pampas foxes breed in the early spring, with the female coming into
heat In thermodynamics, heat is energy in transfer between a thermodynamic system and its surroundings by such mechanisms as thermal conduction, electromagnetic radiation, and friction, which are microscopic in nature, involving sub-atomic, ato ...
just once each year. After a
gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...
period of 55 to 60 days, the mother gives birth to a litter of up to eight kits. The young are born between September and December, and are weaned around two months of age. Females reach sexual maturity in their first year, and animals have lived for up to 14 years in captivity. Pups remain in dens until at least the age of 3 months, when they start hunting with parents. The males bring food to their females who stay at the den with kits.


Threats

The main threats to the Pampas fox comes from humans hunting them for their fur or to prevent them from attacking livestock; they may be affected by the loss of their natural habitat. However, because they remain common in most areas where they have been studied, the Pampas fox is not presently considered a threatened species.


Hybridisation with dog

Crossings between canids of different
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
are extremely rare. In 2021, a female canid with unusual phenotypic characteristics was found in Vacaria,
Rio Grande do Sul Rio Grande do Sul (, ; ; "Great River of the South") is a Federative units of Brazil, state in the South Region, Brazil, southern region of Brazil. It is the Federative units of Brazil#List, fifth-most populous state and the List of Brazilian s ...
, Brazil. DNA analysis indicates that the canid was a hybrid between a Pampas fox and a domestic dog. Dubbed a ' Dogxim' or 'graxorra', this finding is the first documented case of hybridisation detected between these two species.


References


External links


Canid Specialist Group: Pampas Fox
{{Taxonbar, from=Q832036 South American foxes Mammals of Argentina Mammals of Bolivia Carnivorans of Brazil Mammals of Paraguay Mammals of Uruguay Pampas Least concern biota of South America pampas fox