''Lukousaurus'' is an
archosauromorph based on most of a small skull's snout, displaying distinctive
lachrymal horns, found in the
Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
-age
Lower Lufeng Formation,
Yunnan
Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
,
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
and was described by
Chung Chien Young in 1940.
The generic name refers to the
Lugou Bridge, lit. “crossroads”, near
Beijing
Beijing, Chinese postal romanization, previously romanized as Peking, is the capital city of China. With more than 22 million residents, it is the world's List of national capitals by population, most populous national capital city as well as ...
, where the
Sino-Japanese War started.
[C.-C. Young. (1940). Preliminary notes on the Lufeng vertebrate fossils. ''Bulletin of the Geological Society of China'' 20(3-4):235-239
] ''L. yini'' is tentatively classified as a theropod
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
by some allied to ceratosaurs, by others a coelurosaur. Its skull is rather robust for its size though the teeth were described by the author as typically
theropod
Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
an. Whatever ''Lukousaurus'' was, it was definitely an
archosauromorph.
[Knoll F., Rohrberg K. (2012). CT scanning, rapid prototyping and re-examination of a partial skull of a basal crocodylomorph from the Late Triassic of Germany. '']Swiss Journal of Geosciences
Springer Science+Business Media, commonly known as Springer, is a German multinational publishing company of books, e-books and peer-reviewed journals in science, humanities, technical and medical (STM) publishing.
Originally founded in 1842 in ...
'' 105:109–115.
History
At a locality in the town of Huangchiatien (also called Dahungtien) in
Yunnan Province
Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
,
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
during the late 1930s, a partial anterior skull and lower jaws as well as possibly a tooth and humerus found nearby.
The skull was found in the
lower Jurassic
The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic–Jurassic extinction event, 201.3 Ma&nb ...
strata of the Red Beds of the
Lufeng Formation
The Lufeng Formation (formerly Lower Lufeng Series) is a Lower Jurassic sedimentary rock formation found in Yunnan, China. It has two units: the lower Dull Purplish Beds/Shawan Member are of Hettangian age, and Dark Red Beds/Zhangjia'ao Member a ...
, though at the time of its naming in 1940 by
Chung Chien Young the beds were thought to date to the Triassic.
Young noted that the skull is very strange, with morphologies similar to those of not just Coelurosaurs, which he thought the taxon was, but also Prosauropods and Carnosaurs.
The generic name refers to the
Lugou Bridge, lit. “crossroads”, near
Beijing
Beijing, Chinese postal romanization, previously romanized as Peking, is the capital city of China. With more than 22 million residents, it is the world's List of national capitals by population, most populous national capital city as well as ...
, where the
Sino-Japanese War started and a symbol of the Chinese resistance against
Japanese imperialism
The territorial conquests of the Empire of Japan in the Western Pacific Ocean and East Asia began in 1895 with its victory over Qing China in the First Sino-Japanese War. Subsequent victories over the Russian Empire (Russo-Japanese War) and the ...
.
The species name honors the former deputy
Director of the Geological Survey of China, T. H. Yin, who kept work on the survey continuing despite the Japanese invasion.
The holotype specimen is housed within the
Institute of Vertebrate Paleontology and Paleoanthropology
The Institute of Vertebrate Paleontology and Paleoanthropology (IVPP; ) of People's Republic of China, China is a research institution and collections repository for fossils, including many dinosaur and pterosaur specimens (many from the Yixian For ...
in
Beijing, China
Beijing, Chinese postal romanization, previously romanized as Peking, is the capital city of China. With more than 22 million residents, it is the world's List of national capitals by population, most populous national capital city as well as ...
under specimen number IVPP 23.
Simmons, 1965 assigned a distal humerus and co-ossified tibia and fibula to ''Lukousaurus'', though there is no overlap between elements.
Classification
The classification of ''Lukousaurus'' is very uncertain due to the strange characteristics of the type specimen and its referred humerus. ''Lukousaurus'' was first described as a Coelurosaur based on its postorbital and the shaped of the orbit compared to ''
Saltopus
''Saltopus'' ("hopping foot") is a genus of very small bipedal dinosauriform containing the single species ''Saltopus elginensis'' from the late Triassic period of Scotland. It is one of the most famous Elgin Reptiles.
Description
''Saltopu ...
,
Podokesaurus
''Podokesaurus'' is a genus of coelophysoid dinosaur that lived in what is now the eastern United States during the Early Jurassic Period. The first fossil was discovered by the geologist Mignon Talbot near Mount Holyoke, Massachusetts, in 19 ...
,'' and ''
Ammosaurus'', though none of these taxa are now considered Coelurosaurs.
The size and "some characteristics of the skull" were also noted as being similar to ''
Palaeosaurus
''Palaeosaurus'' (or ''Paleosaurus'') is a genus of indeterminate archosaur known from two teeth found in the Bromsgrove Sandstone Formation and also either the Magnesian Conglomerate or the Avon Fissure Fill of Clifton, Bristol, England (or ...
'', though ''Palaeosaurus'' is considered dubious. Dong Zhiming in his book on Chinese dinosaurs assigned ''Lukousaurus'' to Podokesauridae in 1992, a similar conclusion to Young's original classification with ''Podokesaurus,'' though in 1997 Kenneth Carpenter placed it as an indeterminate Theropod''.'' In 2008, Mickey Mortimer believed that ''Lukousaurus'' was either an abelisaurid ceratosaur or a sphenosuchian, but revised her position in 2011 after placing it as a basal pseudosuchian using a phylogenetic matrix developed by Sterling Nesbitt. Knoll ''et al''., 2012 found ''Lukousaurus'' to be similar to
suchian archosaurs in the anatomy of the antorbital fenestra
and Irmis, 2004 stated that ''Lukousaurus'' wasn't a theropod dinosaur or a
dinosauromorph in general.
[Irmis, R. B. (2004). First report of Megapnosaurus (Theropoda: Coelophysoidea) from China. ''PaleoBios'', ''24''(3), 11-18.]
References
{{Taxonbar, from=Q591610
Taxa named by Yang Zhongjian
Fossil taxa described in 1940
Archosauromorpha
Lufeng Formation