''Luchibang'' (meaning "
Lü's heron wing") is an extinct genus of
istiodactylid pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the Order (biology), order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cre ...
discovered in
Inner Mongolia
Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an autonomous region of the People's Republic of China. Its border includes most of the length of China's border with the country of Mongolia. Inner Mongolia also accounts for ...
,
China. The type and only species is ''Luchibang xingzhe''.
Discovery and naming
The
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
, a juvenile specimen, was acquired from illegal fossil dealers who found it before 2000, probably at the village of Liutiaogou, near
Dashuangmiao in
Ningcheng, Inner-Mongolia. Around 2009, David Hone was asked by
Xu Xing to describe the specimen, but publication of the article was for many years delayed due to doubts expressed in the
peer review
Peer review is the evaluation of work by one or more people with similar competencies as the producers of the work ( peers). It functions as a form of self-regulation by qualified members of a profession within the relevant field. Peer revie ...
about its authenticity. ''Luchibang'' was announced in an abstract of the Los Angeles ''Flugsaurier'' conference in 2018, but it was only validly published in 2020.
In 2020, David W.E. Hone, Adam James Fitch, Ma Feimin and Xu Xing named and described the
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
''Luchibang xingzhe''. The generic name ''Luchibang'' is derived from the Mandarin ''lu'', "heron", and ''chibang'', "wing", but is also a reference to the late paleontologist
Lü Junchang. The
specific name ''xingzhe'' means "walker", in reference to its terrestrial capabilities.
In the Hone et al. paper, the section formally describing ''Luchibang'' spells the species name as ''xinzhe''; however, most of the paper uses the name ''xingzhe''. In addition, one figure and the supplemental data contain the name ''Luchibang wuke''. A
corrigendum
An erratum or corrigendum (plurals: errata, corrigenda) (comes from la, errata corrige) is a correction of a published text. As a general rule, publishers issue an erratum for a production error (i.e., an error introduced during the publishing pro ...
to the paper clarifies that the spelling ''xinzhe'' was in error, and that ''wuke'' was the original planned name but lacks a description and is therefore a ''
nomen nudum
In taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published with an adequate des ...
''. Hone et al. formally changed the species name from ''xinzhe'' to the intended ''xingzhe''.
The holotype ELDM 1000 is assumed to have been found in a layer of the
Yixian Formation
The Yixian Formation (; formerly transcribed as Yihsien Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the late Barremian and early Aptian stages of the Early Cretaceous. It is known for its ...
dating from the
Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
. It consists of a nearly complete skeleton with skull, compressed on a single plate. Only the rear of the skull, the front two neck vertebrae, the sternal ribs, the tail, a prepubis, the left wrist and left pteroid are lacking. Pieces of skin are preserved. The skeleton is largely articulated and visible from the underside. It represents a juvenile individual. It is part of the collection of the
Erlianhaote Dinosaur Museum
Erenhot ( mn, ; , commonly shortened to Ereen or Erlian) is a county-level city of the Xilin Gol League, in Inner Mongolia Autonomous Region, located in the Gobi Desert along the Sino-Mongolian border, across from the Mongolian town of Zamy ...
.
The fossil had already been prepared by the traders. Due to differences in colouration and the fact that the proportions of the postcrania, especially the long legs and large feet, resemble those of
Azhdarchoidea
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with ...
, it was suggested in the peer review that the piece might be a chimaera, the head having been added to the rump to increase its value. To check this, Hone and C. Rodgers further prepared the head region but could find no trace of connecting glue. The head seemed to have been an integral part of the piece and therefore was concluded to be authentic.
Description
The holotype individual has a wing span of about two metres. As it is a juvenile, the adult size might have been considerably larger, which would make ''Luchibang'' quite a large istiodactylid.
The describing authors indicated some distinguishing traits. Two of these are
autapomorphies
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to t ...
, unique derived characters. The sternum is large and rectangular with a straight rear edge. The femur equals more than 80% of the ulna length.
Additionally, a unique combination is present of traits that are in themselves not unique. The front of the sternum shows no vertical expansion. In the rear jaws, the teeth are far apart. In top view, the symphysis of the lower jaw dentaries is four times longer than wide. The branches of the lower jaws are long and narrow, twenty times longer than wide in top view.
Palaeobiology
Two specimens of the fish ''
Lycoptera'' were found in association with the holotype; the first, resting between the jaws of the pterosaur, likely represents an animal that died around the same time, while the second was found within or underneath the ribcage of the pterosaur, and "may represent gut contents" according to the authors.
[ However, due to the high level of compression involved in the preservation of the fossil, it is impossible to determine whether the associated fish skeleton truly lies within the ribcage, and thus represents true evidence for its diet, or if it is merely underneath the pterosaur skeleton, a possibility that the authors themselves discuss.
]
References
{{Portal bar, Paleontology, Cretaceous, China
Pteranodontoids
Early Cretaceous pterosaurs of Asia
Fossil taxa described in 2020
Paleontology in Inner Mongolia