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Upland and lowland are conditional descriptions of a
plain In geography, a plain, commonly known as flatland, is a flat expanse of land that generally does not change much in elevation, and is primarily treeless. Plains occur as lowlands along valleys or at the base of mountains, as coastal plains, and ...
based on
elevation The elevation of a geographic location (geography), ''location'' is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational equipotenti ...
above
sea level Mean sea level (MSL, often shortened to sea level) is an mean, average surface level of one or more among Earth's coastal Body of water, bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical ...
. In studies of the
ecology Ecology () is the natural science of the relationships among living organisms and their Natural environment, environment. Ecology considers organisms at the individual, population, community (ecology), community, ecosystem, and biosphere lev ...
of
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mi ...
river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...
s,
habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s are classified as upland or lowland.


Definitions

Upland and lowland are portions of a plain that are conditionally categorized by their elevation above the
sea level Mean sea level (MSL, often shortened to sea level) is an mean, average surface level of one or more among Earth's coastal Body of water, bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical ...
. Lowlands are usually no higher than , while uplands are somewhere around to . On unusual occasions, certain lowlands such as the Caspian Depression lie below sea level. Uplands areas tend to spike into
valley A valley is an elongated low area often running between hills or mountains and typically containing a river or stream running from one end to the other. Most valleys are formed by erosion of the land surface by rivers or streams over ...
s and
mountain A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited summit area, and is usually higher t ...
s, forming
mountain ranges A mountain range or hill range is a series of mountains or hills arranged in a line and connected by high ground. A mountain system or mountain belt is a group of mountain ranges with similarity in form, structure, and alignment that have arise ...
while lowland areas tend to be uniformly flat, although both can vary such as the Mongolian Plateau. Upland habitats are cold, clear and rocky whose rivers are fast-flowing in mountainous areas; lowland habitats are warm with slow-flowing rivers found in relatively flat lowland areas, with water that is frequently colored by
sediment Sediment is a solid material that is transported to a new location where it is deposited. It occurs naturally and, through the processes of weathering and erosion, is broken down and subsequently sediment transport, transported by the action of ...
and organic matter. These classifications overlap with the geological definitions of "upland" and "lowland". In
geology Geology (). is a branch of natural science concerned with the Earth and other astronomical objects, the rocks of which they are composed, and the processes by which they change over time. Modern geology significantly overlaps all other Earth ...
an "upland" is generally considered to be land that is at a higher elevation than the
alluvial plain An alluvial plain is a plain (an essentially flat landform) created by the deposition of sediment over a long period by one or more rivers coming from highland regions, from which alluvial soil forms. A ''floodplain'' is part of the process, bei ...
or stream terrace, which are considered to be "lowlands". The term "bottomland" refers to low-lying alluvial land near a river. Much freshwater fish and invertebrate communities around the world show a pattern of specialization into upland or lowland river habitats. Classifying rivers and streams as upland or lowland is important in freshwater ecology, as the two types of river habitat are very different, and usually support very different populations of fish and invertebrate species.


Uplands

In freshwater ecology, upland rivers and
stream A stream is a continuous body of water, body of surface water Current (stream), flowing within the stream bed, bed and bank (geography), banks of a channel (geography), channel. Depending on its location or certain characteristics, a strea ...
s are the fast-flowing rivers and streams that drain elevated or mountainous country, often onto broad alluvial plains (where they become lowland rivers). However, elevation is not the sole determinant of whether a river is upland or lowland. Arguably the most important determinants are those of
stream power Stream power, originally derived by Ralph Bagnold, R. A. Bagnold in the 1960s, is the amount of energy the water in a river or stream is exerting on the sides and bottom of the river. Stream power is the result of multiplying the density of the wa ...
and stream gradient. Rivers with a course that drops rapidly in elevation will have faster water flow and higher stream power or "force of water". This in turn produces the other characteristics of an upland river—an incised course, a river bed dominated by
bedrock In geology, bedrock is solid rock that lies under loose material ( regolith) within the crust of Earth or another terrestrial planet. Definition Bedrock is the solid rock that underlies looser surface material. An exposed portion of bed ...
and coarse sediments, a riffle and pool structure and cooler water temperatures. Rivers with a course that drops in elevation very slowly will have slower water flow and lower force. This in turn produces the other characteristics of a lowland river—a meandering course lacking
rapids Rapids are sections of a river where the river bed has a relatively steep stream gradient, gradient, causing an increase in water velocity and turbulence. Flow, gradient, constriction, and obstacles are four factors that are needed for a rapid t ...
, a river bed dominated by fine sediments and higher water temperatures. Lowland rivers tend to carry more suspended sediment and organic matter as well, but some lowland rivers have periods of high
water clarity Water clarity is a descriptive term for how deeply visible light penetrates through water. In addition to light penetration, the term water clarity is also often used to describe underwater visibility. Water clarity is one way that humans measure ...
in seasonal low-flow periods. The generally clear, cool, fast-flowing waters and bedrock and coarse sediment beds of upland rivers encourage fish species with limited temperature tolerances, high oxygen needs, strong swimming ability and specialised reproductive strategies to prevent eggs or larvae being swept away. These characteristics also encourage invertebrate species with limited temperature tolerances, high oxygen needs and ecologies revolving around coarse sediments and interstices or "gaps" between those coarse sediments. The term "upland" is also used in wetland ecology, where "upland" plants indicate an area that is not a wetland.


Lowlands

The generally more
turbid Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of both water clarity and wate ...
, warm, slow-flowing waters and fine sediment beds of lowland rivers encourage fish species with broad temperature tolerances and greater tolerances to low oxygen levels, and life history and breeding strategies adapted to these and other traits of lowland rivers. These characteristics also encourage invertebrate species with broad temperature tolerances and greater tolerances to low oxygen levels and ecologies revolving around fine sediments or alternative habitats such as submerged woody debris ("snags") or submergent macrophytes ("water weed").


Lowland alluvial plains

Lowland alluvial plains form when there is deposition of sediment over a long period of time by one or more rivers coming from highland regions, and then are deposited in lowland regions for long periods of time. Examples include American Bottom, a flood plain of the Mississippi River in Southern Illinois, Bois Brule Bottom, and Bottomland hardwood forest a deciduous hardwood forest found in broad lowland floodplains of the United States.


See also

* Freshwater biology *
Highland Highlands or uplands are areas of high elevation such as a mountainous region, elevated mountainous plateau or high hills. Generally, ''upland'' refers to a range of hills, typically from up to , while ''highland'' is usually reserved for range ...
* Mountain river * River reclamation *
Riparian zone A riparian zone or riparian area is the interface between land and a river or stream. In some regions, the terms riparian woodland, riparian forest, riparian buffer zone, riparian corridor, and riparian strip are used to characterize a ripari ...
*
Plateau In geology and physical geography, a plateau (; ; : plateaus or plateaux), also called a high plain or a tableland, is an area of a highland consisting of flat terrain that is raised sharply above the surrounding area on at least one side. ...


References

{{Authority control Freshwater ecology Water and the environment Riparian zone Rivers *