Longan
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''Dimocarpus longan'', commonly known as the longan () and dragon's eye, is a tropical tree species that produces edible fruit. It is one of the better-known tropical members of the soapberry family
Sapindaceae The Sapindaceae are a family (biology), family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1,858 accepted species. Examples include Aesculus, horse chestnut, maples, ackee and lychee. The ...
, to which the
lychee Lychee ( , ; ''Litchi chinensis''; ) is a monotypic taxon and the sole member in the genus ''Litchi'' in the soapberry family, Sapindaceae. There are three distinct subspecies of lychee. The most common is the Indochinese lychee found in So ...
and rambutan also belong. The fruit of the longan is similar to that of the lychee, but is less aromatic in taste. The longan (from Vietnamese ''long nhãn'' or
Cantonese Cantonese is the traditional prestige variety of Yue Chinese, a Sinitic language belonging to the Sino-Tibetan language family. It originated in the city of Guangzhou (formerly known as Canton) and its surrounding Pearl River Delta. While th ...
''lùhng ngáahn'' , literally 'dragon eye'), is so named because the black seed within the shelled fruit creates the appearance of an eyeball. The plant is native to tropical Asia and China.


Description

Depending upon climate and soil type the tree may grow to over in height, but it typically stands in height and the
crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, parti ...
is round. The trunk is thick with corky
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
. The branches are long and thick, typically drooping. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are oblong and blunt-tipped, usually long and wide. The leaves are pinnately compounded and alternate. There are 6 to 9 pairs of leaflets per leaf and the upper surface is wavy and a dark, glossy-green. The longan tree produces light-yellow
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
s at the end of branches. The inflorescence is commonly called a
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
; they can be long, and widely branched. The small
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s have 5 to 6
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s and brownish-yellow
petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...
s. The flower has a two-lobed
pistil Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
and 8
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
. There are three flower types, distributed throughout the panicle; staminate (functionally male), pistillate (functionally female), and hermaphroditic flowers. Flowering occurs as a progression. The fruit are spherical and about wide; they hang in drooping clusters. The shell is tan, thin, and leathery with tiny hairs; when firm, it can be squeezed (as in the cracking of a
sunflower seed A sunflower seed is a seed from a sunflower (''Helianthus annuus''). There are three types of commonly used sunflower seeds: linoleic (most common), high oleic, and sunflower oil seeds. Each variety has its own unique levels of monounsatura ...
) to shell the fruit. The flesh is translucent, and the
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
is large and black with a circular white spot at the base. This gives the illusion of an eye. The flesh has a musky, sweet taste, which can be compared to the flavor of
lychee Lychee ( , ; ''Litchi chinensis''; ) is a monotypic taxon and the sole member in the genus ''Litchi'' in the soapberry family, Sapindaceae. There are three distinct subspecies of lychee. The most common is the Indochinese lychee found in So ...
fruit. The seed is round, hard, and has a lacquered appearance. The longan tree is somewhat sensitive to
frost Frost is a thin layer of ice on a solid surface, which forms from water vapor that deposits onto a freezing surface. Frost forms when the air contains more water vapor than it can normally hold at a specific temperature. The process is simila ...
. While the species prefers temperatures that do not typically fall below , it can withstand brief temperature drops to about . Longan trees prefer sandy soil with mild levels of acidity and
organic matter Organic matter, organic material or natural organic matter is the large source of carbon-based compounds found within natural and engineered, terrestrial, and aquatic environments. It is matter composed of organic compounds that have come fro ...
. Longans usually bear fruit slightly later than lychees.


Taxonomy

The longan is believed to originate from the mountain range between
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
and southern China. Other reported origins include Indonesia, India,
Sri Lanka Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...
,
upper Myanmar Upper Myanmar ( or , also called Upper Burma) is one of two geographic regions in Myanmar, the other being Lower Myanmar. Located in the country's centre and north stretches, Upper Myanmar encompasses six inland states and regions, including M ...
, north Thailand, Kampuchea (more commonly known as Cambodia), north
Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
and
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; , fossilized , also known as Papua or historically ) is the List of islands by area, world's second-largest island, with an area of . Located in Melanesia in the southwestern Pacific Ocean, the island is ...
. Its earliest record of existence draws back to the
Han dynasty The Han dynasty was an Dynasties of China, imperial dynasty of China (202 BC9 AD, 25–220 AD) established by Liu Bang and ruled by the House of Liu. The dynasty was preceded by the short-lived Qin dynasty (221–206 BC ...
in 200 BCE. The emperor had demanded lychee and longan trees to be planted in his palace gardens in
Shaanxi Shaanxi is a Provinces of China, province in north Northwestern China. It borders the province-level divisions of Inner Mongolia to the north; Shanxi and Henan to the east; Hubei, Chongqing, and Sichuan to the south; and Gansu and Ningxia to t ...
, but the plants failed. Four hundred years later, longan trees flourished in other parts of China like
Fujian Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...
and
Guangdong ) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
, where longan production soon became an industry. Later on, due to immigration and the growing demand for nostalgic foods, the longan tree was officially introduced to Australia in the mid-1800s, Thailand in the late-1800s, and Hawaii and Florida in the 1900s. The warm, sandy-soiled conditions allowed for the easy growth of longan trees. This jump-started the longan industry in these locations. Despite its long success in China, the longan is considered to be a relatively new fruit to the world. It has only been acknowledged outside of China in the last 250 years. The first European acknowledgment of the fruit was recorded by
João de Loureiro João de Loureiro (1717, Lisbon – 18 October 1791) was a Portuguese people, Portuguese Jesuit missionary and botanist. Biography After receiving admission to the Jesuit Order, João de Loureiro served as a missionary in Goa, capital of Port ...
, a Portuguese Jesuit botanist, in 1790. The first entry resides in his collection of works, ''Flora Cochinchinensis''.


Subspecies

Plants of the World Online Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew. History Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...
lists: * ''D. longan'' var. ''echinatus'' Leenhouts (Borneo, Philippines) * ''D. longan'' var. ''longetiolatus'' Leenhouts (Viet Nam) * ''D. longan'' subsp. ''malesianus'' Leenh. (widespread SE Asia) * ''D. longan'' var. ''obtusus'' (Pierre) Leenh. (Indo-China)


Conservation

The wild longan population have been decimated considerably by large-scale logging in the past, and the species used to be listed as Vulnerable on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
. If left alone, longan tree stumps will resprout and the listing was upgraded to
Near Threatened A near-threatened species is a species which has been Conservation status, categorized as "Near Threatened" (NT) by the International Union for Conservation of Nature (IUCN) as that may be vulnerable to Endangered species, endangerment in the ne ...
in 1998. Recent field data are inadequate for a contemporary IUCN assessment.


Diseases

Plant based diseases can affect both longan fruits and their trees, and the severity of these diseases can range from harmless cosmetic damage to rendering to the fruit inedible. The most prevalent disease among longan plants is witch's broom, which can be found in all major longan-producing Asian territories, including China, Thailand, and Vietnam. Witch's broom deforms longan skin, and at times causes the plant to prematurely drop its fruit, similar to the ''
Phytophthora palmivora ''Phytophthora palmivora'' is an oomycete that causes bud-rot of Arecaceae, palms, fruit-rot or kole-roga of coconut and areca nut. These are among the most serious diseases caused by fungi and Mold (fungus), moulds in South India. Outbreaks occu ...
''. Another common disease that longan trees can carry is the aptly named ''longan decline'', which is largely prevalent in Thailand, with reports finding that it could affect up to 40% of longan trees alone. Affected trees are more vulnerable to common tree pests and algae, and often bear low-quality fruit unworthy of yield. ''Algal spot'' is another plant disease that can affect longan plants and trees. Common among tropical fruits, the disease mainly takes form as red-orange algae that can appear on a fruit-bearing tree's leaves or branches.''Algal spot'' on longan plants, like many other tropical fruits, is caused by '' Cephaleuros virescens''. An oomycete disease that causes
blight Blight is a specific symptom affecting plants in response to infection by a pathogenic organism. Description Blight is a rapid and complete chlorosis, browning, then death of plant tissues such as leaves, branches, twigs, or floral organs. A ...
on leaves and foliage of a plant and affects the related lychee, ''Phytophthora palmivora'', can also appear on both longan plants and fruit, particularly in the Thailand region. When affecting longan, it can create brown spots on the fruit in an erratic fashion, and can also cause longan to drop prematurely from the plant. Early symptoms can also include a dark
necrosis Necrosis () is a form of cell injury which results in the premature death of cells in living tissue by autolysis. The term "necrosis" came about in the mid-19th century and is commonly attributed to German pathologist Rudolf Virchow, who i ...
on the plant itself. Stem-end rot is a disease common amongst litchi and longan, and causes browning and rot on the stem of the fruit. Longan also suffer from various decay-accelerating fungi. An oomycete disease that affects the related lychee, '' Phytophthora litchii'', also afflicts ''D. longan''.


Cultivation

Currently, longan crops are grown in southern China,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
, northern Thailand, Malaysia, Indonesia, Cambodia, Laos, Vietnam, India, Sri Lanka, the
Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
,
Bangladesh Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...
,
Mauritius Mauritius, officially the Republic of Mauritius, is an island country in the Indian Ocean, about off the southeastern coast of East Africa, east of Madagascar. It includes the main island (also called Mauritius), as well as Rodrigues, Ag ...
, the United States, and Australia.


Growth

Longan, like its sister fruit lychee, thrives in humid areas or places with high rainfall, and can grow on most types of soil that does not induce issues with water drainage. Ample temperatures are also instrumental in longan growth: while longan can resist small stretches of cool temperatures, they can be damaged or killed in longer stretches of temperatures as high as −2 degrees Celsius. Younger plants tend to be more vulnerable to the cold than those more mature.


Harvest

During harvest, pickers must climb ladders to carefully remove branches of fruit from longan trees. Longan fruit remain fresher if still attached to the branch, so efforts are made to prevent the fruit from detaching too early. Mechanical picking would damage the delicate skin of the fruit, so the preferred method is to harvest by hand. Knives and scissors are the most commonly used tools. Fruit is picked early in the day to minimize water loss and to prevent high heat exposure, which would be damaging. The fruit is then placed into either plastic crates or bamboo baskets and taken to packaging houses, where the fruit undergo a series of checks for quality. The packaging houses are well-ventilated and shaded to prevent further decay. The process of checking and sorting are performed by workers instead of machinery. Any fruit that is split, under-ripe, or decaying is disposed of. The remaining healthy fruit is then prepared and shipped to markets. Many companies add preservatives to canned longan. Regulations control the preserving process. The only known preservative added to canned longan is
sulfur dioxide Sulfur dioxide (IUPAC-recommended spelling) or sulphur dioxide (traditional Commonwealth English) is the chemical compound with the formula . It is a colorless gas with a pungent smell that is responsible for the odor of burnt matches. It is r ...
, to prevent discoloration. Fresh longan that is shipped worldwide is exposed to sulfur
fumigation Fumigation is a method of pest control or the removal of harmful microorganisms by completely filling an area with gaseous pesticides, or fumigants, to suffocate or poison the pests within. It is used to control pests in buildings (structural ...
. Tests have shown that sulfur residues remain on the fruit skin, branches, and leaves for a few weeks. This violates many countries' limits on fumigation residue, and efforts have been made to reduce this amount.


Distribution

Longan is found commonly in most of Asia, primarily in mainland China, Taiwan, Vietnam and Thailand. China, the main longan-producing country in the world, produced about of longan in 2015–2017, accounting for 70% of the world's longan production and more than 50% of the world's longan plots. Vietnam and Thailand produced around , respectively. Like Vietnam, Thailand's economy relies heavily on the cultivation and shipments of longan as well as lychee. This increase in the production of longan reflects recent interest in exotic fruits in other parts of the world. However, the majority of the demand comes from Asian communities in North America, Europe and Australia.


Yield

While longan yields average out to 2 to 5 tonnes per hectare, there have been observed yields of up to 19.5 tonnes per ha in
Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...
. Advancements in
selective breeding Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant m ...
have allowed scientists to find a strain of longan containing a "high proportion of aborted seeds" at the end of a thirty-year breeding program in 2001. Studies in 2015 that aimed to aid longan breeding efforts discovered that −20 degrees Celsius is the optimal temperature for long-term storage of longan pollen, a key ingredient in enabling longan breeding programs.


Uses


Nutrition

Raw longan fruit is 83% water, 15%
carbohydrate A carbohydrate () is a biomolecule composed of carbon (C), hydrogen (H), and oxygen (O) atoms. The typical hydrogen-to-oxygen atomic ratio is 2:1, analogous to that of water, and is represented by the empirical formula (where ''m'' and ''n'' ...
s, 1%
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
, and contains negligible
fat In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food. The term often refers specif ...
(table). In a reference amount, raw longan supplies 60
calorie The calorie is a unit of energy that originated from the caloric theory of heat. The large calorie, food calorie, dietary calorie, kilocalorie, or kilogram calorie is defined as the amount of heat needed to raise the temperature of one liter o ...
s of food energy, 93% of the
Daily Value In the U.S. and Canada, the Reference Daily Intake (RDI) is used in nutrition labeling on food and dietary supplement products to indicate the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97â ...
(DV) of
vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits, berries and vegetables. It is also a generic prescription medication and in some countries is sold as a non-prescription di ...
, 11% DV of
riboflavin Riboflavin, also known as vitamin B2, is a vitamin found in food and sold as a dietary supplement. It is essential to the formation of two major coenzymes, flavin mononucleotide and flavin adenine dinucleotide. These coenzymes are involved in ...
, and no other
micronutrient Micronutrients are essential chemicals required by organisms in small quantities to perform various biogeochemical processes and regulate physiological functions of cells and organs. By enabling these processes, micronutrients support the heal ...
s in appreciable quantities (table).


Culinary

The fruit is sweet, juicy, and succulent in superior agricultural varieties. The seed and the peel are not consumed. Apart from being eaten raw like other fruits, longan fruit is also often used in Asian soups, snacks, desserts, and sweet-and-sour foods, either fresh or dried, and sometimes preserved and canned in syrup. The taste is different from lychees; while longan has a drier sweetness similar to dates, lychees are often messily juicy with a more tropical, grape-like sour sweetness. Dried longan are often used in
Chinese cuisine Chinese cuisine comprises cuisines originating from Greater China, China, as well as from Overseas Chinese, Chinese people from other parts of the world. Because of the Chinese diaspora and the historical power of the country, Chinese cuisine ...
and Chinese sweet dessert soups. In
Chinese food therapy Chinese food therapy (, also called nutrition therapy and dietary therapy) is a mode of dieting rooted in Chinese beliefs concerning the effects of food on the human organism, and centered on concepts such as seasonal eating and in moderation. ...
and
herbal medicine Herbal medicine (also called herbalism, phytomedicine or phytotherapy) is the study of pharmacognosy and the use of medicinal plants, which are a basis of traditional medicine. Scientific evidence for the effectiveness of many herbal treatments ...
, it is believed to have an effect on relaxation. In contrast with the fresh fruit, which is juicy and white, the flesh of dried longans is dark brown to almost black. Once fermented, it can be made into longan wine.


See also

*''
Lansium parasiticum ''Epicharis parasitica'', commonly known as yellow mahogany, is a species of tree in the family Meliaceae; it grows primarily in tropical rainforests and is native to Taiwan, parts of Malesia, Papuasia, and northeast Queensland. Description ''E ...
'', the langsat or lanzones *'' Talisia esculenta'', a visually similar fruit from South America


References


External links


Longan Production in Asia
from the
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations; . (FAO) is a specialized agency of the United Nations that leads international efforts to defeat hunger and improve nutrition and food security. Its Latin motto, , translates ...
of the United Nations {{Authority control Articles containing video clips Cantonese cuisine Chinese fruit
longan ''Dimocarpus longan'', commonly known as the longan () and dragon's eye, is a tropical tree species that produces edible fruit. It is one of the better-known tropical members of the soapberry family Sapindaceae, to which the lychee and rambu ...
Edible fruits Flora of tropical Asia Fruits originating in East Asia Han dynasty Flora of the Indomalayan realm Plants described in 1790 Traditional Chinese medicine Trees of China Tropical fruit