This is a list of animals for which there is documented evidence of homosexual behavior. These animals have been observed practicing homosexual

courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage or committed romantic, ''de facto'' relationship. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marri ...

sexual behavior Human sexual activity, human sexual practice or human sexual behaviour is the manner in which humans experience and express their sexuality. People engage in a variety of sexual acts, ranging from activities done alone (e.g., masturbation) t ...

affection Affection or fondness is a "disposition or state of mind or body" commonly linked to a feeling or type of love. It has led to multiple branches in philosophy and psychology that discuss emotion, disease, influence, and state of being. Often, " ...

pair bond In biology, a pair bond is the strong affinity that develops in some species between a mating pair, often leading to the production and rearing of young and potentially a lifelong bond. Pair-bonding is a term coined in the 1940s that is frequently ...

ing, or

parenting Parenting or child rearing promotes and supports the physical, cognitive, social, emotional, and educational development from infancy to adulthood. Parenting refers to the intricacies of raising a child and not exclusively for a biologica ...

. Bruce Bagemihl writes that the presence of same-sex sexual behavior was not officially observed on a large scale until the 1990s due to possible

observer bias Observer bias is one of the types of detection bias and is defined as any kind of systematic divergence from accurate facts during observation and the recording of data and information in studies. The definition can be further expanded upon to inc ...

caused by social attitudes towards

LGBT LGBTQ people are individuals who are lesbian, gay, bisexual, transgender, queer, or questioning. Many variants of the initialism are used; LGBTQIA+ people incorporates intersex, asexual, aromantic, agender, and other individuals. The gro ...

people, which made homosexuality in animals a

taboo A taboo is a social group's ban, prohibition or avoidance of something (usually an utterance or behavior) based on the group's sense that it is excessively repulsive, offensive, sacred or allowed only for certain people.''Encyclopædia Britannica ...

subject. News-medical.net (2006) He devotes three chapters, "Two Hundred Years at Looking at Homosexual Wildlife", "Explaining (Away) Animal Homosexuality", and "Not For Breeding Only" in his 1999 book ''Biological Exuberance'' to the "documentation of systematic

prejudice Prejudice can be an affect (psychology), affective feeling towards a person based on their perceived In-group and out-group, social group membership. The word is often used to refer to a preconceived (usually unfavourable) evaluation or classifi ...

s" where he notes "the ''present ignorance'' of

biology Biology is the scientific study of life and living organisms. It is a broad natural science that encompasses a wide range of fields and unifying principles that explain the structure, function, growth, History of life, origin, evolution, and ...

lies precisely in its single-minded attempt to find reproductive (or other) "explanations" for homosexuality, transgender, and non-procreative and alternative

heterosexual Heterosexuality is romantic attraction, sexual attraction, or sexual behavior between people of the opposite sex or gender. As a sexual orientation, heterosexuality is "an enduring pattern of emotional, romantic, and/or sexual attractions ...

ities.Bagemihl (1999) page 213 Petter Bøckman, academic adviser for the '' Against Nature?'' exhibit, stated " ny researchers have described homosexuality as something altogether different from sex. They must realise that animals can have sex with who they will, when they will and without consideration to a researcher's

ethical Ethics is the philosophical study of moral phenomena. Also called moral philosophy, it investigates normative questions about what people ought to do or which behavior is morally right. Its main branches include normative ethics, applied e ...

principle A principle may relate to a fundamental truth or proposition that serves as the foundation for a system of beliefs or behavior or a chain of reasoning. They provide a guide for behavior or evaluation. A principle can make values explicit, so t ...

s". Homosexual behavior is found amongst social

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s and

mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...

s, particularly the sea mammals and the

primates Primates is an order of mammals, which is further divided into the strepsirrhines, which include lemurs, galagos, and lorisids; and the haplorhines, which include tarsiers and simians ( monkeys and apes). Primates arose 74–63 ...

. Animal sexual behavior takes many different forms, even within the same

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

and the motivations for and implications of their behaviors have yet to be fully understood. Bagemihl's research shows that homosexual behavior, not necessarily sexual activity, has been documented in about 500 species as of 1999, ranging from

to gut worms.Harrold (1999) Homosexuality in animals is controversial with some

social conservatives Social conservatism is a political philosophy and a variety of conservatism which places emphasis on traditional social structures over social pluralism. Social conservatives organize in favor of duty, traditional values and social instit ...

because it asserts the naturalness of homosexuality in humans, while others counter that it has no implications and is nonsensical to equate animal behavior to

morality Morality () is the categorization of intentions, Decision-making, decisions and Social actions, actions into those that are ''proper'', or ''right'', and those that are ''improper'', or ''wrong''. Morality can be a body of standards or principle ...

. Animal preference and motivation is inferred from behavior, thus homosexual behavior has been given a number of terms over the years. Modern research applies the term ''homosexuality'' to all sexual behavior ( copulation, genital stimulation, mating games and sexual

display behavior Display may refer to: Technology * Display device, output device for presenting information, including: ** Electronic visual display, output device to present information for visual or tactile reception *** Cathode-ray tube (CRT), that uses an el ...

) between animals of the same sex. This is a list of some

mammals A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle e ...

that have been recorded engaging in homosexual behavior, which is part of a larger list of animals displaying homosexual behavior including

birds Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

insects Insects (from Latin ') are hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed ...

fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...

, etc.

Selected images

Image:Macaques à bonnet petits.jpg, Male

bonnet macaque The bonnet macaque (''Macaca radiata''), also known as zati,Chambers English Dictionary is a species of macaque endemism, endemic to southern India. Its distribution is limited by the Indian Ocean on three sides and the Godavari River, Godavari a ...

s, similar to the youthful ones pictured, "give each other hand-jobs and sometimes eat the resulting

semen Semen, also known as seminal fluid, is a bodily fluid that contains spermatozoon, spermatozoa which is secreted by the male gonads (sexual glands) and other sexual organs of male or hermaphrodite, hermaphroditic animals. In humans and placen ...

" although using "hand-job" can be seen as overly

anthropomorphic Anthropomorphism is the attribution of human traits, emotions, or intentions to non-human entities. It is considered to be an innate tendency of human psychology. Personification is the related attribution of human form and characteristics to ...

. Image:Uganda-Kob.jpg, Ugandan kob - "Female kob perform

oral sex Oral sex, sometimes referred to as oral intercourse, is sexual activity involving the stimulation of the genitalia of a person by another person using the mouth (including the lips, tongue, or teeth). Cunnilingus is oral sex performed on the vu ...

on each other and even stroke each other's vulvas with their forelegs. They may exhibit

urolagnia Urolagnia, also known as urophilia, is a paraphilia in which sexual excitement is associated with urine or urination. Etymologically, the term comes from the Greek , meaning 'urine', and , meaning 'lust'. A golden shower is slang for the practic ...

during sex, one female will

urinate Urination is the release of urine from the bladder through the urethra in placental mammals, or through the cloaca in other vertebrates. It is the urinary system's form of excretion. It is also known medically as micturition, voiding, ure ...

while the other sticks her nose in the stream."Imaginova (2007e) Image:Spotted hyena2.jpg, "Elevated levels of

testosterone Testosterone is the primary male sex hormone and androgen in Male, males. In humans, testosterone plays a key role in the development of Male reproductive system, male reproductive tissues such as testicles and prostate, as well as promoting se ...

in utero" increases aggressiveness and both male and female

spotted hyena The spotted hyena (''Crocuta crocuta''), also known as the laughing hyena, is a hyena species, currently classed as the sole extant member of the genus ''Crocuta'', native to sub-Saharan Africa. It is listed as being of least concern by the IUC ...

s mount submissive same-sex members who likely have lower levels of testosterone from their mothers.Forger (1998)Holekamp (2003)Wilson (Sexing the Hyena) Image:Graywhale MMC.jpg, In "slip-and-slide" orgies, groups of male grey whales, roll in the

ocean The ocean is the body of salt water that covers approximately 70.8% of Earth. The ocean is conventionally divided into large bodies of water, which are also referred to as ''oceans'' (the Pacific, Atlantic, Indian Ocean, Indian, Southern Ocean ...

rubbing their bellies against each other so that their

genitals A sex organ, also known as a reproductive organ, is a part of an organism that is involved in sexual reproduction. Sex organs constitute the primary sex characteristics of an organism. Sex organs are responsible for producing and transporting ...

are touching.Bagemihl (1999) page 339Imaginova (2007h)

List

American bison The American bison (''Bison bison''; : ''bison''), commonly known as the American buffalo, or simply buffalo (not to be confused with Bubalina, true buffalo), is a species of bison that is endemic species, endemic (or native) to North America. ...

Bagemihl (1999) page 413 *

Antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...

Asian elephant The Asian elephant (''Elephas maximus''), also known as the Asiatic elephant, is the only living ''Elephas'' species. It is the largest living land animal in Asia and the second largest living Elephantidae, elephantid in the world. It is char ...

Bagemihl (1999) page 432 * Asiatic mouflonBagemihl (1999) pages 405, 690 *

Atlantic spotted dolphin The Atlantic spotted dolphin (''Stenella frontalis'') is a dolphin found in warm temperate and tropical waters of the Atlantic Ocean. Older members of the species have a very distinctive spotted coloration all over their bodies. Taxonomy The Atl ...

Australian sea lion The Australian sea lion (''Neophoca cinerea''), also known as the Australian sea-lion or Australian sealion, is a species of sea lion that is the only endemic pinniped in Australia. It is currently Monotypic taxon, monotypic in the genus ''Neopho ...

Bagemihl (1999) page 367 *

Barasingha The barasingha (''Rucervus duvaucelii''), sometimes barasinghe, also known as the swamp deer, is a deer species distributed in the Indian subcontinent. Populations in northern and central India are fragmented, and two isolated populations occu ...

Bagemihl (1999) page 378 *

Barbary sheep The Barbary sheep (''Ammotragus lervia''), also known as aoudad (pronounced �ɑʊdæd, is a species of caprine native to rocky mountains in North Africa and parts of West Africa. While this is the only species in genus ''Ammotragus'', six sub ...

Bagemihl (1999) page 405 *

Beluga Beluga may refer to: Animals *Beluga (sturgeon) * Beluga whale Vehicles * Airbus Beluga, a large transport airplane * Airbus BelugaXL, a larger transport airplane * Beluga-class submarine, a class of Russian SSA diesel-electric submarine * U ...

Bharal The bharal (''Pseudois nayaur''), also called the blue sheep, is a Caprinae, caprine native to the high Himalayas. It is the monotypic taxon, only member of the genus ''Pseudois.'' It occurs in Pakistan, India, Nepal, Bhutan, Myanmar, and in Ch ...

Bagemihl (1999) pages 209, 408, 690 *

Bighorn sheep The bighorn sheep (''Ovis canadensis'') is a species of Ovis, sheep native to North America. It is named for its large Horn (anatomy), horns. A pair of horns may weigh up to ; the sheep typically weigh up to . Recent genetic testing indicates th ...

* Black bear *

Blackbuck The blackbuck (''Antilope cervicapra''), also known as the Indian antelope, is a medium-sized antelope native to India and Nepal. It inhabits grassy plains and lightly forested areas with perennial water sources. It stands up to high at the sh ...

Bagemihl (1999) page 402 * Black-footed rock wallabyBagemihl (1999) page 449 *

Black-tailed deer Black-tailed deer or blacktail deer occupy coastal regions of western North America. There are two subspecies, the Columbian black-tailed deer (''Odocoileus hemionus columbianus'') which ranges from the Pacific Northwest of the United States and ...

Bonin flying fox The Bonin flying fox (''Pteropus pselaphon''), also known as Bonin fruit bat, is a species of flying fox in the family Pteropodidae. It is endemic to four islands (Chichijima, Hahajima, North Iwo Jima, and South Iwo Jima) of the Bonin Island ...

Bonnet macaque The bonnet macaque (''Macaca radiata''), also known as zati,Chambers English Dictionary is a species of macaque endemism, endemic to southern India. Its distribution is limited by the Indian Ocean on three sides and the Godavari River, Godavari a ...

Bagemihl (1999) page 316 *

Bonobo The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee (less often the dwarf chimpanzee or gracile chimpanzee), is an endangered great ape and one of the two species making up the genus ''Pan (genus), Pan'' (the other bei ...

de Waal (2001) *

Bottlenose dolphin The bottlenose dolphin is a toothed whale in the genus ''Tursiops''. They are common, cosmopolitan members of the family Delphinidae, the family of oceanic dolphins. Molecular studies show the genus contains three species: the common bot ...

Bowhead whale The bowhead whale (''Balaena mysticetus''), sometimes called the Greenland right whale, Arctic whale, and polar whale, is a species of baleen whale belonging to the family Balaenidae and is the only living representative of the genus '' Balaena' ...

* Brazilian guinea pigBagemihl (1999) page 467 * Bridled dolphin *

Brown bear The brown bear (''Ursus arctos'') is a large bear native to Eurasia and North America. Of the land carnivorans, it is rivaled in size only by its closest relative, the polar bear, which is much less variable in size and slightly bigger on av ...

Bagemihl (1999) page 441 *

Brown capuchin The tufted capuchin (''Sapajus apella''), also known as brown capuchin, black-capped capuchin, or pin monkey, is a New World monkey, New World primate from South America and the Caribbean islands of Trinidad and Margarita Island, Margarita. As t ...

Bagemihl (1999) page 334 * Brown long-eared batBagemihl (1999) page 473 *

Brown rat The brown rat (''Rattus norvegicus''), also known as the common rat, street rat, sewer rat, wharf rat, Hanover rat, Norway rat and Norwegian rat, is a widespread species of common rat. One of the largest Muroidea, muroids, it is a brown or grey ...

Bagemihl (1999) page 469 * Buffalo *

Caribou The reindeer or caribou (''Rangifer tarandus'') is a species of deer with circumpolar distribution, native to Arctic, subarctic, tundra, boreal, and mountainous regions of Northern Europe, Siberia, and North America. It is the only represe ...

Bagemihl (1999) pages 388,389 * Cat (domestic)Bagemihl (1999) pages 81, 88 * Cattle (domestic)Bagemihl (1999) pages 81, 82, 89 *

Chacma baboon The chacma baboon (''Papio ursinus''), also known as the Cape baboon, is, like all other baboons, from the Old World monkey family. It is one of the largest of all monkeys. Located primarily in southern Africa, the chacma baboon has a wide vari ...

Poiani (2010) page 52 *

Cheetah The cheetah (''Acinonyx jubatus'') is a large Felidae, cat and the Fastest animals, fastest land animal. It has a tawny to creamy white or pale buff fur that is marked with evenly spaced, solid black spots. The head is small and rounded, wit ...

Chimpanzee The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...

Bagemihl (1999) pages 276–279
Excerpt from the book:
Common Chimpanzees engage in full mouth-to-mouth contact ..Oral sex of various kinds also occurs in a number of species for examplecunnilingus in Common Chimpanzees ..In ..Common Chimpanzees, individuals often rub their anal and genital regions together ..Other ..forms of "manual" stimulation include ..anal stimulation and penetration with fingers by male Common Chimpanzees. *

Chital The chital or cheetal (''Axis axis''; ), also called spotted deer, chital deer and axis deer, is a deer species native to the Indian subcontinent. It was first described by Johann Christian Polycarp Erxleben in 1777. A moderate-sized deer, mal ...

Poiani (2010) page 51 *

Collared peccary The collared peccary (''Dicotyles tajacu'') is a peccary, a species of artiodactyl (even-toed) mammal in the family Peccary, Tayassuidae found in North America, North, Central America, Central, and South America. It is the only member of the gen ...

Bagemihl (1999) pages 422–425 *

Commerson's dolphin Commerson's dolphin (''Cephalorhynchus commersonii''), also referred to by the common names jacobita, skunk dolphin, piebald dolphin, panda dolphin, or tonina overa (in South America), is a small oceanic dolphin of the genus ''Cephalorhynchus''. ...

Common brushtail possum The common brushtail possum (''Trichosurus vulpecula'', from the Ancient Greek, Greek for "furry tailed" and the Latin for "little fox", previously in the genus ''Phalangista'') is a nocturnal, semiarboreal marsupial of the family Phalangeridae ...

Bagemihl (1999) page 457 *

Common dolphin The common dolphin (''Delphinus delphis'') is the most abundant cetacean in the world, with a global population of about six million. Despite this fact and its vernacular name, the common dolphin is not thought of as the archetypal dolphin, wit ...

* Common marmoset * Common pipistrelleBagemihl (1999) page 475 * Common tree shrewBagemihl (1999) page 471 * Common wallaroo *

Cotton-top tamarin The cotton-top tamarin (''Saguinus oedipus'') is a small New World monkey weighing less than . This New World monkey can live up to 24 years, but most of them die by 13 years. One of the smallest primates, the cotton-top tamarin is easily reco ...

Bagemihl (1999) page 333 *

Crab-eating macaque The crab-eating macaque (''Macaca fascicularis''), also known as the long-tailed macaque or cynomolgus macaque, is a cercopithecine primate native to Southeast Asia. As a synanthropic species, the crab-eating macaque thrives near human settlem ...

* Crested black macaque * Dall's sheep * Daubenton's bat * Dog (domestic)Bagemihl (1999) pages 310, 314 *

Donkey The donkey or ass is a domesticated equine. It derives from the African wild ass, ''Equus africanus'', and may be classified either as a subspecies thereof, ''Equus africanus asinus'', or as a separate species, ''Equus asinus''. It was domes ...

Poiani (2010) page 170 * Doria's tree kangaroo *

Dugong The dugong (; ''Dugong dugon'') is a marine mammal. It is one of four living species of the order Sirenia, which also includes three species of manatees. It is the only living representative of the once-diverse family Dugongidae; its closest ...

Bagemihl (1999) page 376 * Dwarf cavy * Dwarf mongooseBagemihl (1999) pages 447–448 *

Eastern cottontail rabbit The eastern cottontail (''Sylvilagus floridanus'') is a New World cottontail rabbit, a member of the family Leporidae. It is the most common rabbit species in North America. Distribution The eastern cottontail can be found in meadows and shrubby ...

Eastern grey kangaroo The eastern grey kangaroo (''Macropus giganteus'': gigantic large-foot; also great grey kangaroo or forester kangaroo) is a marsupial found in the eastern third of Australia, with a population of several million. Although a large ''M. giganteus ...

* Elk *

European bison The European bison (: bison) (''Bison bonasus'') or the European wood bison, also known as the wisent ( or ), the zubr (), or sometimes colloquially as the European buffalo, is a European species of bison. It is one of two extant species of bi ...

* European polecat *

Fallow deer Fallow deer is the common name for species of deer in the genus ''Dama'' of subfamily Cervinae. There are two living species, the European fallow deer (''Dama dama''), native to Europe and Anatolia, and the Persian fallow deer (''Dama mesopotamic ...

False killer whale The false killer whale (''Pseudorca crassidens'') is a species of oceanic dolphin that is the only extant representative of the genus ''Pseudorca''. It is found in oceans worldwide but mainly in tropical regions. It was first species descriptio ...

* Fat-tailed dunnartBagemihl (1999) pages 458–460 *

Fin whale The fin whale (''Balaenoptera physalus''), also known as the finback whale or common rorqual, is a species of baleen whale and the second-longest cetacean after the blue whale. The biggest individual reportedly measured in length, wi ...

* FoxBagemihl (1999) pages 218, 231, 317 * François' langur *

Gazelle A gazelle is one of many antelope species in the genus ''Gazella'' . There are also seven species included in two further genera; '' Eudorcas'' and '' Nanger'', which were formerly considered subgenera of ''Gazella''. A third former subgenus, ' ...

Bagemihl (1999) page 391 *

Gelada baboon The gelada (''Theropithecus gelada'', , ), sometimes called the bleeding-heart monkey or the gelada baboon, is a species of Old World monkey found only in the Ethiopian Highlands, living at elevations of above sea level. It is the only living me ...

Bagemihl (1999) pages 324–330 * Goat (domestic) * Golden monkeyBagemihl (1999) pages 299–301 *

Giraffe The giraffe is a large Fauna of Africa, African even-toed ungulate, hoofed mammal belonging to the genus ''Giraffa.'' It is the Largest mammals#Even-toed Ungulates (Artiodactyla), tallest living terrestrial animal and the largest ruminant on ...

Gorilla Gorillas are primarily herbivorous, terrestrial great apes that inhabit the tropical forests of equatorial Africa. The genus ''Gorilla'' is divided into two species: the eastern gorilla and the western gorilla, and either four or five su ...

Bagemihl (1999) pages 280–284 *

Grant's gazelle Grant's gazelle (''Nanger granti'') is a relatively large species of gazelle antelope, distributed from northern Tanzania to South Sudan and Ethiopia, and from the Kenyan coast to Lake Victoria. Its Swahili name is ''swala granti''. It was named ...

* Grey-headed flying fox * Grey seal * Grey squirrelBagemihl (1999) pages 461–464 * Grey whale *

Grey wolf The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, including the dog and dingo, though gr ...

Bagemihl (1999) pages 231, 436–440 *

Grizzly bear The grizzly bear (''Ursus arctos horribilis''), also known as the North American brown bear or simply grizzly, is a population or subspecies of the brown bear inhabiting North America. In addition to the mainland grizzly (''Ursus arctos horr ...

* Guinea pig (domestic) *

Hamadryas baboon The hamadryas baboon (''Papio hamadryas'' ; gawina;Aerts 2019 , Ar Robbaḥ) is a species of baboon within the Old World monkey family. It is the northernmost of all the baboons, being native to the Horn of Africa and the southwestern region o ...

* Hamster (domestic) * Hanuman langurBagemihl (1999) pages 293–298 *

Harbor porpoise The harbour porpoise (''Phocoena phocoena'') is one of eight extant species of porpoise. It is one of the smallest species of cetacean. As its name implies, it stays close to coastal areas or river estuaries, and as such, is the most familiar ...

Bagemihl (1999) page 347 * Harbor seal *

Himalayan tahr The Himalayan tahr (''Hemitragus jemlahicus'') is a large even-toed ungulate native to the Himalayas in southern Tibet, northern India, western Bhutan and Nepal. It is listed as Near Threatened on the IUCN Red List, as the population is declini ...

Bagemihl (1999) page 412 *

Hoary marmot The hoary marmot (''Marmota caligata'') is a species of marmot that inhabits the mountains of northwest North America. Hoary marmots live near the tree line on slopes with Graminoid, grasses and forbs to eat and rocky areas for cover. It is the ...

Bagemihl (1999) page 465-466 * Horse (domestic)Bagemihl (1999) pages 81, 165, 205, 226, 231 *

Human Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...

(see

Human sexual behavior Human sexual activity, human sexual practice or human sexual behaviour is the manner in which humans experience and express their sexuality. People engage in a variety of sexual acts, ranging from activities done alone (e.g., masturbation) t ...

) *

Humpback whale The humpback whale (''Megaptera novaeangliae'') is a species of baleen whale. It is a rorqual (a member of the family Balaenopteridae) and is the monotypic taxon, only species in the genus ''Megaptera''. Adults range in length from and weigh u ...

* Indian fruit bat * Indian muntjacBagemihl (1999) page 386 *

Indian rhinoceros The Indian rhinoceros (''Rhinoceros unicornis''), also known as the greater one-horned rhinoceros, great Indian rhinoceros or Indian rhino, is a species of rhinoceros found in the Indian subcontinent. It is the second largest living rhinocer ...

Bagemihl (1999) page 430 *

Japanese macaque The Japanese macaque (''Macaca fuscata''), also known as the snow monkey, is a terrestrial Old World monkey species that is native to Japan. Colloquially, they are referred to as "snow monkeys" because some live in areas where snow covers the g ...

Javelina Peccaries (also javelinas or skunk pigs) are pig-like ungulates of the family Tayassuidae (New World pigs). They are found throughout Central and South America, Trinidad in the Caribbean, and in the southwestern area of North America. Peccari ...

Bagemihl (1999) page 422-425 *

Kangaroo rat Kangaroo rats, small mostly nocturnal rodents of genus ''Dipodomys'', are native to arid areas of western North America. The common name derives from their bipedal form. They hop in a manner similar to the much larger kangaroo, but developed thi ...

Killer whale The orca (''Orcinus orca''), or killer whale, is a toothed whale and the largest member of the oceanic dolphin family. The only extant species in the genus '' Orcinus'', it is recognizable by its black-and-white-patterned body. A cosmopolit ...

Koala The koala (''Phascolarctos cinereus''), sometimes inaccurately called the koala bear, is an arboreal herbivorous marsupial native to Australia. It is the only Extant taxon, extant representative of the Family (biology), family ''Phascolar ...

Bagemihl (1999) page 455-457 * KobBagemihl (1999) page 397-401 * Larga seal * Least chipmunk * Lechwe * Lesser bushbabyBagemihl (1999) page 336-338 *

Lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'', native to Sub-Saharan Africa and India. It has a muscular, broad-chested body (biology), body; a short, rounded head; round ears; and a dark, hairy tuft at the ...

Lion-tailed macaque The lion-tailed macaque (''Macaca silenus''), also known as the wanderoo, is an Old World monkey endemic to the Western Ghats of South India. Characteristics Lion-tailed macaques are covered in black fur, and have a striking gray or silver ...

* Lion tamarin *

Little brown bat The little brown bat or little brown myotis (''Myotis lucifugus'') is an endangered species of mouse-eared bat, mouse-eared microbat found in North America. It has a small body size and glossy brown fur. It is similar in appearance to several ...

* Livingstone's fruit bat * Long-eared hedgehogBagemihl (1999) pages 470–472 * Long-footed tree shrew *

Macaque The macaques () constitute a genus (''Macaca'') of gregarious Old World monkeys of the subfamily Cercopithecinae. The 23 species of macaques inhabit ranges throughout Asia, North Africa, and Europe (in Gibraltar). Macaques are principally f ...

* MarkhorBagemihl (1999) page 409 *

Marten A marten is a weasel-like mammal in the genus ''Martes'' within the subfamily Guloninae, in the family Mustelidae. They have bushy tails and large paws with partially retractile claws. The fur varies from yellowish to dark brown, depending on ...

Bagemihl (1999) page 448 *

Masked palm civet The masked palm civet (''Paguma larvata''), also called the gem-faced civet or Himalayan palm civet, is a viverrid species native to the Indian subcontinent and Southeast Asia. It has been listed as least concern on the IUCN Red List since 2008 ...

* MocoBagemihl (1999) pages 109, 469 * Mohol galago * Moor macaque *

Moose The moose (: 'moose'; used in North America) or elk (: 'elk' or 'elks'; used in Eurasia) (''Alces alces'') is the world's tallest, largest and heaviest extant species of deer and the only species in the genus ''Alces''. It is also the tal ...

Bagemihl (1999) pages 387–390 *

Mountain goat The mountain goat (''Oreamnos americanus''), also known as the Rocky Mountain goat, is a cloven-footed mammal that is endemic to the remote and rugged mountainous areas of western North America. A subalpine to truly alpine species, it is a s ...

* Mountain tree shrew *

Mountain zebra The mountain zebra (''Equus zebra'') is a zebra species in the family Equidae, native to southwestern Africa. There are two subspecies, the Cape mountain zebra (''E. z. zebra'') found in South Africa and Hartmann's mountain zebra (''E. z. hartma ...

Bagemihl (1999) pages 418–421 * Mouse (domestic)Bagemihl (1999) pages 663, 693, 714 * Moustached tamarin *

Mule deer The mule deer (''Odocoileus hemionus'') is a deer indigenous to western North America; it is named for its ears, which are large like those of the mule. Two subspecies of mule deer are grouped into the black-tailed deer. Unlike the related whit ...

* Musk-oxBagemihl (1999) pages 410–413 *

Natterer's bat Natterer's bat (''Myotis nattereri'') is a European Vespertilionidae, vespertilionid bat with pale wings. It has brown fur tending to greyish-white on its underside. It is found across most of the continent of Europe, parts of the Near East and No ...

* New Zealand sea lion * Nilgiri langur * Noctule *

North American porcupine The North American porcupine (''Erethizon dorsatum''), also known as the Canadian porcupine, is a large quill-covered rodent in the New World porcupine family. It is the second largest rodent in North America after the North American beaver (''Ca ...

Bagemihl (1999) page 472 *

Northern elephant seal The northern elephant seal (''Mirounga angustirostris'') is one of two species of elephant seal (the other is the southern elephant seal). It is a member of the family Phocidae (true seals). Elephant seals derive their name from their great s ...

Northern fur seal The northern fur seal (''Callorhinus ursinus'') is an eared seal found along the north Pacific Ocean, the Bering Sea, and the Sea of Okhotsk. It is the largest member of the fur seal subfamily (Arctocephalinae) and the only living species in the ...

* Northern quoll *

Olympic marmot The Olympic marmot (''Marmota olympus'') is a rodent in the squirrel family, Sciuridae. It occurs only in the U.S. state of Washington, at the middle elevations of the Olympic Peninsula. The closest relatives of this species are the hoary mar ...

Bagemihl (1999) pages 465–466 *

Orangutan Orangutans are great apes native to the rainforests of Indonesia and Malaysia. They are now found only in parts of Borneo and Sumatra, but during the Pleistocene they ranged throughout Southeast Asia and South China. Classified in the genus ...

Bagemihl (1999) pages 284–288 * Pacific striped dolphin * Patas monkeyBagemihl (1999) page 328 * Pere David's deer * Pig (domestic)Bagemihl (1999) pages 81, 82, 225–226, 232–240 * Pig-tailed macaque *

Plains zebra The plains zebra (''Equus quagga'', formerly ''Equus burchellii'') is the most common and geographically widespread species of zebra. Its range is fragmented, but spans much of southern and eastern Africa south of the Sahara. Six or seven subspec ...

Bagemihl (1999) page 418-421 *

Polar bear The polar bear (''Ursus maritimus'') is a large bear native to the Arctic and nearby areas. It is closely related to the brown bear, and the two species can Hybrid (biology), interbreed. The polar bear is the largest extant species of bear ...

* Pretty-faced wallaby *

Proboscis monkey The proboscis monkey or long-nosed monkey (''Nasalis larvatus'') is an arboreal Old World monkey with an unusually large nose (or proboscis), a reddish-brown skin color and a long tail. It is endemic to the southeast Asian island of Borneo an ...

Pronghorn The pronghorn (, ) (''Antilocapra americana'') is a species of artiodactyl (even-toed, hoofed) mammal indigenous to interior western and central North America. Though not an antelope, it is known colloquially in North America as the American ante ...

Bagemihl (1999) pages 394–396 * Pudú * PukuBagemihl (1999) pages 397–401 * QuokkaBagemihl (1999) page 451 *

Rabbit Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...

Bagemihl (1999) page 81 *

Raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the North American, northern or common raccoon (also spelled racoon) to distinguish it from Procyonina, other species of raccoon, is a mammal native to North America. It is the largest ...

Raccoon dog ''Nyctereutes'' (Greek: ''nyx, nykt-'' "night" + ''ereutēs'' "wanderer") is a genus of canid which includes only two extant species, both known as raccoon dogs: the common raccoon dog (''Nyctereutes procyonoides'') and the Japanese raccoon do ...

Bagemihl (1999) page 440 *

Red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or Hart (deer), hart, and a female is called a doe or hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Ir ...

Red fox The red fox (''Vulpes vulpes'') is the largest of the true foxes and one of the most widely distributed members of the order Carnivora, being present across the entire Northern Hemisphere including most of North America, Europe and Asia, plus ...

Bagemihl (1999) pages 437–441 *

Red kangaroo The red kangaroo (''Osphranter rufus'') is the largest of all kangaroos, the largest terrestrial mammal native to Australia, and the Largest mammals#Marsupials (Marsupialia), largest extant marsupial. It is found across mainland Australia, exce ...

* Red-necked wallaby *

Red squirrel The red squirrel (''Sciurus vulgaris''), also called Eurasian red squirrel, is a species of tree squirrel in the genus ''Sciurus''. It is an arboreal and primarily herbivorous rodent and common throughout Eurasia. Taxonomy There have been ...

Reeves's muntjac Reeves's muntjac (''Muntiacus reevesi''), also known as the Chinese muntjac, is a species of muntjac found widely in south-eastern China (from Gansu to Yunnan) and Taiwan. It has also been introduced in the United Kingdom, Ireland, the Netherla ...

Reindeer The reindeer or caribou (''Rangifer tarandus'') is a species of deer with circumpolar distribution, native to Arctic, subarctic, tundra, taiga, boreal, and mountainous regions of Northern Europe, Siberia, and North America. It is the only re ...

Rhesus macaque The rhesus macaque (''Macaca mulatta''), colloquially rhesus monkey, is a species of Old World monkey. There are between six and nine recognised subspecies split between two groups, the Chinese-derived and the Indian-derived. Generally brown or g ...

Right whale Right whales are three species of large baleen whales of the genus ''Eubalaena'': the North Atlantic right whale (''E. glacialis''), the North Pacific right whale (''E. japonica'') and the southern right whale (''E. australis''). They are class ...

* Rock cavy * Rodrigues fruit bat * Roe deer * Rufous bettongBagemihl (1999) pages 453–455 * Rufous-naped tamarin * Rufous rat kangaroo * Saddle-back tamarin * Savanna baboon *

Sea otter The sea otter (''Enhydra lutris'') is a marine mammal native to the coasts of the northern and eastern Pacific Ocean, North Pacific Ocean. Adult sea otters typically weigh between , making them the heaviest members of ...

Bagemihl (1999) pages 364–365 * Serotine bat * Sheep (domestic)Roselli (2004), Vol. 145, No. 2, pages 478–483 *

Siamang The siamang (, ; ''Symphalangus syndactylus'') is an endangered arboreal, black-furred gibbon native to the forests of Indonesia, Malaysia, and Thailand. The largest of the gibbons, the siamang can be twice the size of other gibbons, reaching i ...

Bagemihl (1999) pages 289–292 *

Sika deer The sika deer (''Cervus nippon''), also known as the northern spotted deer or the Japanese deer, is a species of deer native to much of East Asia and introduced to other parts of the world. Previously found from northern Vietnam in the south t ...

* Slender tree shrew * Snub-nosed monkey * Sooty mangabey *

Sperm whale The sperm whale or cachalot (''Physeter macrocephalus'') is the largest of the toothed whales and the largest toothed predator. It is the only living member of the Genus (biology), genus ''Physeter'' and one of three extant species in the s ...

Spider monkey Spider monkeys are New World monkeys belonging to the genus ''Ateles'', part of the subfamily Atelinae, family Atelidae. Like other atelines, they are found in tropical forests of Central and South America, from southern Mexico to Brazil. The g ...

* Spinifex hopping mouse * Spinner dolphin *

Spotted hyena The spotted hyena (''Crocuta crocuta''), also known as the laughing hyena, is a hyena species, currently classed as the sole extant member of the genus ''Crocuta'', native to sub-Saharan Africa. It is listed as being of least concern by the IUC ...

Forger (6 December 1998), Volume 375, Issue 2, Pages 333 – 343 * Spotted seal * Squirrel monkeyBagemihl (1999) pages 330–335 * Striped dolphin * Stuart's marsupial mouseBagemihl (1999) page 459 * Stumptail macaque * Swamp deer * Swamp wallaby * Takhi * Talapoin * Tammar wallaby *

Tasmanian devil The Tasmanian devil (''Sarcophilus harrisii''; palawa kani: ''purinina'') is a carnivorous marsupial of the family Dasyuridae. It was formerly present across mainland Australia, but became extinct there around 3,500 years ago; it is now con ...

* Tibetan macaque * Tasmanian rat kangaroo * Thinhorn sheep *

Thomson's gazelle Thomson's gazelle (''Eudorcas thomsonii'') is one of the best known species of gazelles. It is named after explorer Joseph Thomson (explorer), Joseph Thomson and is sometimes referred to as a "tommie". It is considered by some to be a subspecies o ...

Tiger The tiger (''Panthera tigris'') is a large Felidae, cat and a member of the genus ''Panthera'' native to Asia. It has a powerful, muscular body with a large head and paws, a long tail and orange fur with black, mostly vertical stripes. It is ...

Sommer (2006) * Tonkean macaque * TucuxiBagemihl (1999) page 340 * UrialBagemihl (1999) pages 405–409 * Ursine colobus *

Vampire bat Vampire bats, members of the subfamily Desmodontinae, are Phyllostomidae, leaf-nosed bats currently found in Central and South America. Their food source is the blood of other animals, a dietary trait called hematophagy. Three extant bat species ...

* Verreaux's sifakaBagemihl (1999) pages 366–368 * Vervet * VicunaBagemihl (1999) pages 425–426 *

Walrus The walrus (''Odobenus rosmarus'') is a large pinniped marine mammal with discontinuous distribution about the North Pole in the Arctic Ocean and subarctic seas of the Northern Hemisphere. It is the only extant species in the family Odobeni ...

Bagemihl (1999) pages 370–374Imaginova (2007g) * WapitiBagemihl (1999) page 231 * WarthogBagemihl (1999) page 421 *

Waterbuck The waterbuck (''Kobus ellipsiprymnus'') is a large antelope found widely in sub-Saharan Africa. It is placed in the genus ''Kobus (antelope), Kobus'' of the family Bovidae. It was first Scientific description, described by Irish naturalist Will ...

Bagemihl (1999) pages 397–400 *

Water buffalo The water buffalo (''Bubalus bubalis''), also called domestic water buffalo, Asian water buffalo and Asiatic water buffalo, is a large bovid originating in the Indian subcontinent and Southeast Asia. Today, it is also kept in Italy, the Balkans ...

* Weeper capuchin *

Western grey kangaroo The western grey kangaroo (''Macropus fuliginosus''), also referred to as a western grey giant kangaroo, black-faced kangaroo, mallee kangaroo, sooty kangaroo and (when referring to the Kangaroo Island subspecies) Kangaroo Island grey kangaroo, i ...

West Indian manatee The West Indian manatee (''Trichechus manatus''), also known as the North American manatee, is a large, aquatic mammal native to warm coastal areas of the Caribbean, from the Eastern United States to northern Brazil. Living alone or in herds, it ...

Bagemihl (1999) pages 374–377 * Whiptail wallaby * White-faced capuchin * White-fronted capuchin * White-handed gibbonBagemihl (1999) pages 288–290 * White-lipped peccaryBagemihl (1999) page 424 *

White-tailed deer The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...

* Wild cavy *

Wild goat The wild goat (''Capra aegagrus'') is a wild goat species, inhabiting forests, shrublands and rocky areas ranging from Turkey and the Caucasus in the west to Turkmenistan, Afghanistan and Pakistan in the east. It has been listed as near threaten ...

Yellow-bellied marmot The yellow-bellied marmot (''Marmota flaviventer''), also known as the rock chuck, is a large, stout-bodied ground squirrel in the marmot genus. It is one of fourteen species of marmots, and is native to mountainous and semi-arid regions of south ...

Poiani (2010) page 50 * Yellow-footed rock wallaby * Yellow-toothed cavy

Bibliography

References

{{DEFAULTSORT:Mammals displaying homosexual behavior

Mammals A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle e ...

Homosexual Homosexuality is romantic attraction, sexual attraction, or sexual behavior between people of the same sex or gender. As a sexual orientation, homosexuality is "an enduring pattern of emotional, romantic, and/or sexual attractions" exc ...

Mammalian sexuality

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See also

Bibliography

References