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''Limnonectes'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
fork-tongued frog The frog family Dicroglossidae occurs in tropical and subtropical regions of Asia and Africa, with most genera and species being found in Asia. The common name of the family is fork-tongued frogs. The Dicroglossidae were previously considered ...
s of about 75 known species, but new ones are still being described occasionally. They are collectively known as fanged frogs because they tend to have unusually large teeth, which are small or absent in other frogs.


Habitat

These frogs are found throughout East and Southeast Asia, most commonly near forest streams. Multiple species of ''Limnonectes'' may occupy the same area in harmony. Large-bodied species cluster around fast rivers, while smaller ones live among leaf-litter or on stream banks. The Indonesian island of Sulawesi is home to at least 15 species of this frog, only four of which have been formally described.


Lifecycle

Tadpole A tadpole is the larval stage in the biological life cycle of an amphibian. Most tadpoles are fully aquatic, though some species of amphibians have tadpoles that are terrestrial. Tadpoles have some fish-like features that may not be found ...
s of this genus have
adapted In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the po ...
to a variety of conditions. Most species (e.g.
Blyth's river frog Blyth's river frog (''Limnonectes blythii''), also known as Blyth's frog, giant Asian river frog or (ambiguously) giant frog, is a species of frog in the family Dicroglossidae found from Myanmar through western Thailand and the Malay Peninsula (M ...
''L. blythii'' or the fanged river frog ''L. macrodon'') develop normally, with free-swimming tadpoles that eat food. The tadpoles of the
corrugated frog ''Limnonectes khasianus'', commonly known as the corrugated frog, rivulet frog, or sometimes (ambiguously) called "flat-headed frog", is a species of frog in the family Dicroglossidae. It is found in Brunei, India, Indonesia, Malaysia, Myanmar, ...
(''L. laticeps'') are free-swimming but endotrophic, meaning they do not eat but live on stored
yolk Among animals which produce eggs, the yolk (; also known as the vitellus) is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example bec ...
until metamorphosis into frogs. Before, '' L. limborgi'' was assumed to have direct development (eggs hatching as tiny, full-formed frogs), but more careful observations have showed it has free-swimming but endotrophic larvae; this probably applies to the closely related '' L. hascheanus'', too. '' L. larvaepartus'' is the only known species of frog that gives live birth to tadpoles. Parental care is performed by males.


Species

* ''
Limnonectes acanthi ''Limnonectes acanthi'' is a species of frog in the family Dicroglossidae. It is endemic to the Philippines. Its natural habitats are subtropical or tropical dry forests, subtropical or tropical moist lowland forests, subtropical or tropical mois ...
'' (Taylor, 1923) – Busuanga wart frog * '' Limnonectes arathooni'' (Smith, 1927) – Djikoro wart frog; (Endangered) * '' Limnonectes asperatus'' * ''
Limnonectes bannaensis ''Limnonectes bannaensis'' (vernacular name: Banna large-headed frog) is a species of frogs in the family Dicroglossidae. It is found in southern China (southern and western Yunnan, southwestern and southeastern Guangxi, and western and central ...
'' Ye, Fei, Xie & Jiang, 2007 * '' Limnonectes beloncioi'' Herr, Vallejos, Meneses, Abraham, Otterholt, Siler, Rico & Brown, 2021
Mindoro fanged frog Mindoro is the seventh largest and eighth-most populous island in the Philippines. With a total land area of 10,571 km2 ( 4,082 sq.mi ) and has a population of 1,408,454 as of 2020 census. It is located off the southwestern coast of Luz ...
* '' Limnonectes blythii'' –
Blyth's river frog Blyth's river frog (''Limnonectes blythii''), also known as Blyth's frog, giant Asian river frog or (ambiguously) giant frog, is a species of frog in the family Dicroglossidae found from Myanmar through western Thailand and the Malay Peninsula (M ...
, giant Asian river frog * '' Limnonectes cintalubang'' (Matsui, Nishikawa & Eto, 2014) * '' Limnonectes coffeatus'' Phimmachak, Sivongxay, Seateun, Yodthong, Rujirawan, Neang, Aowphol, and Stuart, 2018 * '' Limnonectes conspicillatus'' (Günther, 1872) * '' Limnonectes dabanus'' * ''
Limnonectes dammermani ''Limnonectes dammermani'', Dammerman's wart frog, is a species of frogs in the family Dicroglossidae endemic to the Lesser Sunda Islands of Indonesia, where it can be found on Flores Flores is one of the Lesser Sunda Islands, a group of i ...
'' – Dammerman's wart frog * ''
Limnonectes deinodon ''Limnonectes deinodon'' is a species of fanged frogs in the family Dicroglossidae. It is endemic to peninsular Malaysia and likely also southern Thailand. It was previously confused with ''Limnonectes laticeps'' and ''Limnonectes khasianus''. R ...
'' Dehling, 2014 * '' Limnonectes diuatus'' – eastern Mindanao frog, Tagibo wart frog * '' Limnonectes doriae'' –
Burmese wart frog ''Limnonectes doriae'' (common name: Burmese wart frog, Doria's frog, or red stream frog) is a species of frog in the family Dicroglossidae found in the Southeast Asia. Etymology The specific name ''doriae'' commemorates the Italian naturalis ...
, Doria's frog, or red stream frog * ''
Limnonectes fastigatus ''Limnonectes'' is a genus of fork-tongued frogs of about 75 known species, but new ones are still being described occasionally. They are collectively known as fanged frogs because they tend to have unusually large teeth, which are small or absen ...
'' Stuart, Schoen, Nelson, Maher, Neang, Rowley & McLeod, 2020 * '' Limnonectes ferneri'' * ''
Limnonectes finchi ''Limnonectes finchi'', Finch's wart frog, is a species of frog in the family Dicroglossidae endemic to Sabah, Malaysia, but it might well occur in adjacent Kalimantan. Its natural habitats are tropical moist lowland forests. The main potential ...
'' –
Finch's wart frog ''Limnonectes finchi'', Finch's wart frog, is a species of frog in the family Dicroglossidae endemic to Sabah, Malaysia, but it might well occur in adjacent Kalimantan. Its natural habitats are tropical moist lowland forests. The main potential ...
* '' Limnonectes fragilis'' * ''
Limnonectes fujianensis ''Limnonectes fujianensis'' (Fujian large-headed frog) is a species of frog in the family Dicroglossidae. It is most closely related to, and formerly confused with ''Limnonectes kuhlii''. Its name refers to the type locality in Fujian province of ...
'' * '' Limnonectes grunniens'' * '' Limnonectes gyldenstolpei'' * '' Limnonectes hascheanus'' (sometimes separated in ''Taylorana'') * '' Limnonectes heinrichi'' * '' Limnonectes hikidai'' Matsui & Nishikawa, 2014 * ''
Limnonectes ibanorum ''Limnonectes ibanorum'' is a species of frog in the family Dicroglossidae. It is endemic to Borneo. Its natural habitats are subtropical or tropical moist lowland forests and rivers. It is becoming rare due to habitat loss. References ...
'' * ''
Limnonectes ingeri ''Limnonectes ingeri'' (common names: Inger's wart frog, greater swamp frog) is a species of frog in the family Dicroglossidae. The species is native to the island of Borneo and associated islets. Etymology The specific name, ''ingeri'', is ...
'' * ''
Limnonectes isanensis ''Limnonectes isanensis'' is a species of fanged frogs in the family Dicroglossidae. It was discovered in Phu Luang Wildlife Sanctuary, Loei Province, Thailand in 2012. It belongs to the '' Limnonectes kuhlii'' species complex.McLeod, Kelly ...
'' McLeod, Kelly, and Barley, 2012 * ''
Limnonectes jarujini ''Limnonectes jarujini'' is a species of frog in the family Dicroglossidae, first described from near Kaeng Krachan Dam The Kaeng Krachan Dam ( th, เขื่อนแก่งกระจาน, , ) is a multi-purpose hydroelectric dam in ...
'' Matsui ''et al.'', 2010 * ''
Limnonectes kadarsani ''Limnonectes kadarsani'' is a species of frog in the family Dicroglossidae. It is endemic to Indonesia where it is found in the Lesser Sunda Islands. Specifically, it has been recorded from Lombok, Sumbawa, Flores, and Adonara. Its natural habi ...
'' * ''
Limnonectes kenepaiensis ''Limnonectes kenepaiensis'' is a species of frog in the family Dicroglossidae. It is endemic to western Borneo and occurs in both Kalimantan (Indonesia) and Sarawak (Malaysia), and perhaps in Sabah (Malaysia). Common name Kenepai wart frog has b ...
'' * ''
Limnonectes khammonensis ''Limnonectes khammonensis'' is a species of frog in the family Dicroglossidae. Endemic to Laos and Vietnam, its status is insufficiently known. References

Limnonectes, khammonensis Amphibians of Laos Amphibians of Vietnam Taxonomy arti ...
'' * ''
Limnonectes khasianus ''Limnonectes khasianus'', commonly known as the corrugated frog, rivulet frog, or sometimes (ambiguously) called "flat-headed frog", is a species of frog in the family Dicroglossidae. It is found in Brunei, India, Indonesia, Malaysia, Myanmar, ...
'' * '' Limnonectes kiziriani'' Pham, Le, Ngo, Ziegler, and Nguyen, 2018 * '' Limnonectes kohchangae'' * '' Limnonectes kong'' Dehling and Dehling, 2017 * ''
Limnonectes kuhlii Kuhl's creek frog or large-headed frog (''Limnonectes kuhlii'') is a species of frog in the family Dicroglossidae. Species complex ''L. kuhlii'' was once believed to have a broad distribution in Northeast India ( Assam) and Southeast Asia, bu ...
'' – Kuhl's Creek frog, large-headed frog * ''
Limnonectes larvaepartus ''Limnonectes larvaepartus'' is a species of fanged frog in the family Dicroglossidae endemic to northern and western Sulawesi, Indonesia. It is unique in that it has internal fertilization and gives live birth to tadpoles. Other frog spe ...
'' * '' Limnonectes lauhachindai'' (Aowphol, Rujirawan, Taksintum, Chuaynkern & Stuart. 2015) * ''
Limnonectes leporinus The giant river frog (''Limnonectes leporinus'') is a species of frog in the family Dicroglossidae. It is endemic to Borneo, and found in Brunei, Kalimantan (Indonesia), and Sabah and Sarawak (Malaysia). Giant river frogs inhabit the banks of sm ...
'' –
giant river frog The giant river frog (''Limnonectes leporinus'') is a species of frog in the family Dicroglossidae. It is endemic to Borneo, and found in Brunei, Kalimantan (Indonesia), and Sabah and Sarawak (Malaysia). Giant river frogs inhabit the banks of sm ...
* '' Limnonectes leytensis'' – small disked frog, swamp frog * ''
Limnonectes limborgi ''Limnonectes limborgi'' is a species of frog in the Dicroglossidae. It is found in Burma, Thailand, Cambodia, Laos, and Vietnam; it might well occur in Northeast India and Yunnan, China. It is a small frog, males being and females snout-vent ...
'' * ''
Limnonectes liui ''Limnonectes liui'' is a species of frog in the family Dicroglossidae. It is found in Yunnan, China, but it likely occurs also in nearby Laos and Myanmar. The taxonomic placement of this species has been much debated and varies between sources. ...
'' (Yang, 1983) * '' Limnonectes longchuanensis'' (Suwannapoom, Yuan, Sullivan & McLeod, 2016) * '' Limnonectes macrocephalus'' – Luzon fanged frog * ''
Limnonectes macrodon The fanged river frog, Javan giant frog, Malaya wart frog, or stone creek frog (''Limnonectes macrodon'') is a species of frog in the family Dicroglossidae endemic to Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It ...
'' (Duméril & Bibron, 1841)fanged river frog, Javan giant frog, Malaya wart frog, stone creek frog * ''
Limnonectes macrognathus ''Limnonectes macrognathus'' is a species of frog in the family Dicroglossidae. It is found in Malaysia, Myanmar and Thailand. Its natural habitats are subtropical or tropical moist lowland forests, rivers and intermittent rivers. Its status is i ...
'' * ''
Limnonectes magnus The giant Philippine frog, large swamp frog, or Mindanao fanged frog (''Limnonectes magnus'') is a species of frog in the family Dicroglossidae. It is endemic to the Philippines. Its natural habitats are tropical moist lowland forests, subt ...
'' (Stejneger, 1910)giant Philippine frog, large swamp frog, Mindanao fanged frog * ''
Limnonectes malesianus The Malesian frog, Malaysian river frog, Malaysian peat frog, or peat swamp frog (''Limnonectes malesianus'') is a species of frog in the family Dicroglossidae. It is found on the Malay Peninsula (including extreme southern peninsular Thailand and ...
'' –
Malesian frog The Malesian frog, Malaysian river frog, Malaysian peat frog, or peat swamp frog (''Limnonectes malesianus'') is a species of frog in the family Dicroglossidae. It is found on the Malay Peninsula (including extreme southern peninsular Thailand ...
, peat swamp frog * ''
Limnonectes mawlyndipi ''Limnonectes mawlyndipi'' is a species of frog in the family Dicroglossidae. It is endemic to India. Its natural habitats are subtropical or tropical moist montane forest, rivers, freshwater marshes, and intermittent freshwater marshes. ...
'' * ''
Limnonectes megastomias ''Limnonectes megastomias'' is a species of frogs with fangs that was discovered in Thailand in 2008. The frog eats birds and insects. The male frogs use their fangs to attack other males in combat. The species have also been known to eat other ...
'' McLeod, 2008 * '' Limnonectes micrixalus'' * ''
Limnonectes microdiscus ''Limnonectes microdiscus'' is a species of frog in the family Dicroglossidae. It is endemic to Indonesia and occurs in Java and southern Sumatra. Its natural habitats are subtropical or tropical moist lowland forest, subtropical or tropical mo ...
'' * ''
Limnonectes microtympanum ''Limnonectes microtympanum'' (common name: Sulawesi wart frog) is a species of frog in the family Dicroglossidae. It is endemic to Sulawesi, Indonesia, where it is only known from the Moncong Lompobatang Moncong Lompobattang is a mountain in ...
'' * '' Limnonectes mocquardi'' (Mocquard, 1890) * '' Limnonectes modestus'' * ''
Limnonectes namiyei ''Limnonectes namiyei'' is a species of frog in the family Dicroglossidae. It is endemic to Okinawa Island, Japan. It is named after Motoyoshi Namiye, a Japanese naturalist and herpetologist. Its common name is Okinawa wart frog or Namiye's frog ...
'' –
Namiye's frog ''Limnonectes namiyei'' is a species of frog in the family Dicroglossidae. It is endemic to Okinawa Island, Japan. It is named after Motoyoshi Namiye, a Japanese naturalist and herpetologist. Its common name is Okinawa wart frog or Namiye's frog; ...
* ''
Limnonectes nguyenorum ''Limnonectes nguyenorum'' is a species of fanged frog in the family Dicroglossidae. It is endemic to northern Vietnam and only known from the area of its type locality in Vi Xuyen District, northwestern Ha Giang Province, northern Vietnam. It i ...
'' * ''
Limnonectes nitidus ''Limnonectes nitidus'' (common name: Tanah Rata wart frog) is a species of frog in the family Dicroglossidae. It is endemic to Peninsular Malaysia where it is only known from the Cameron Highlands and Fraser's Hill, both in Pahang state. ''Li ...
'' * '' Limnonectes palavanensis'' * ''
Limnonectes paramacrodon ''Limnonectes paramacrodon'' (commonly known as the masked swamp frog) is a species of frog in the family Dicroglossidae. It is found in Malay Peninsula The Malay Peninsula (Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainlan ...
'' * ''
Limnonectes parvus The Philippine small-disked frog (''Limnonectes parvus'') is a species of frog in the family Dicroglossidae. It is endemic to the Philippines. Its natural habitats are subtropical or tropical moist lowland forest Tropical and subtropical ...
'' – Philippine small-disked frog * '' Limnonectes plicatellus'' * '' Limnonectes poilani'' * ''
Limnonectes quangninhensis ''Limnonectes quangninhensis'' (Quangninh Wart Frog; Vietnamese: Ếch nhẽo Quảng Ninh) is a species of fanged frog in the family Dicroglossidae. Its type locality is Quảng Sơn Commune, Hải Hà District, Quảng Ninh Province, Vie ...
'' Pham, Le, Nguyen, Ziegler, Wu, and Nguyen, 2017 * '' Limnonectes rhacoda'' * ''
Limnonectes savan ''Limnonectes'' is a genus of fork-tongued frogs of about 75 known species, but new ones are still being described occasionally. They are collectively known as fanged frogs because they tend to have unusually large teeth, which are small or abse ...
'' Phimmachak, Richards, Sivongxay, Seateun, Chuaynkern, Makchai, Som & Stuart, 2019 * ''
Limnonectes selatan ''Limnonectes selatan'' is a species of fanged frogs in the family Dicroglossidae. It is endemic to peninsular Malaysia, in the states of Pahang and Selangor. References

* Matsui, Belabut & Ahmad, 2014 : ''Two new species of fanged frogs fr ...
'' Matsui, Belabut, and Ahmad, 2014 * ''
Limnonectes shompenorum ''Limnonectes shompenorum'' (common name: Shompen frog) is a species of frog in the family Dicroglossidae. It is found in Great Nicobar Island ( India), its type locality, and in western Sumatra ( Indonesia). It is similar to ''Limnonectes m ...
'' * ''
Limnonectes sinuatodorsalis ''Limnonectes sinuatodorsalis'' is a species of fanged frog in the family Dicroglossidae. It is endemic to Borneo, where it was found in the border region of East Kalimantan and Sarawak Sarawak (; ) is a state of Malaysia. The largest among ...
'' Matsui, 2015 * ''
Limnonectes sisikdagu ''Limnonectes sisikdagu'' is a species of fanged frogs in the family Dicroglossidae. It is endemic to West Sumatra, Indonesia, where its holotype was found near Solok. It is part of the ''Limnonectes kuhlii'' species complex. References

* M ...
'' McLeod, Horner, Husted, Barley, and Iskandar, 2011 * '' Limnonectes taylori'' Matsui ''et al.'', 2010 * ''
Limnonectes timorensis ''Limnonectes timorensis'' (common name: Timor river frog and Timor wart frog) is a species of frogs in the family Dicroglossidae. It is endemic to the island of Timor at the eastern end of the Lesser Sunda Islands. In Timor-Leste East Ti ...
'' (Smith, 1927) * ''
Limnonectes tweediei ''Limnonectes tweediei'' (common name: Tweedie's wart frog) is a species of frogs in the family Dicroglossidae. It is found in Sumatra ( Indonesia) and the Malay Peninsula The Malay Peninsula (Malay: ''Semenanjung Tanah Melayu'') is a pe ...
'' * ''
Limnonectes utara ''Limnonectes utara'' is a species of fanged frogs in the family Dicroglossidae. It can be found in Malaysia (in the states of Perak and Terengganu) and Thailand. References * Matsui, Belabut & Ahmad, 2014 : ''Two new species of fanged frogs fr ...
'' Matsui, Belabut, and Ahmad, 2014 * ''
Limnonectes visayanus The giant Visayan frog (''Limnonectes visayanus'') is a species of frog in the family Dicroglossidae. It is endemic to the Philippines, and is known from Masbate, Cebu, Negros, Guimaras, Panay, and Siquijor islands. Its natural habitats are subt ...
'' –
giant Visayan frog The giant Visayan frog (''Limnonectes visayanus'') is a species of frog in the family Dicroglossidae. It is endemic to the Philippines, and is known from Masbate, Cebu, Negros, Guimaras, Panay, and Siquijor islands. Its natural habitats are su ...
* '' Limnonectes woodworthi'' – Woodworth's frog


Phylogeny


Pyron & Wiens (2011)

The following phylogeny of ''Limnonectes'' is from Pyron & Wiens (2011). 35 species are included. ''Limnonectes'' is a
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
of ''
Nanorana ''Nanorana'' is a genus of dicroglossid frogs. They are found in Asia, from the Himalayan region of northern Pakistan and northern India, Nepal, and western China east to montane southern China and southeast to Myanmar, Thailand, Laos, and ...
''.


Aowphol, et al. (2015)

The following ''Limnonectes'' phylogeny is from Aowphol, et al. (2015). 20 species are included.


McLeod, et al. (2015)

Below is a phylogeny of species within the '' L. kuhlii''
species complex In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
(McLeod, et al. 2015). '' Limnonectes longchuanensis'', '' Limnonectes hikidai'', and '' Limnonectes cintalubang''Matsui, Nishikawa, and Eto, 2014, Raffles Bull. Zool., Singapore, 62: 681. Holotype: KUHE 47859, by original designation. Type locality: "Ranchan, Serian, Samarahan Division, Sarawak, East Malaysia (01° 08′ 30″ N, 110° 34′ 57″ E, 64 m asl)". http://zoobank.org/urn:lsid:zoobank.org:pub:9C21B7C4-27AD-4103-89C0-513D2E80106C are also part of the ''L. kuhlii''
species complex In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
.


References

{{Taxonbar, from=Q2288011 Dicroglossidae Amphibians of Asia Amphibian genera Taxa named by Leopold Fitzinger