''Limaysaurus'' ("Limay lizard") is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
represented by a single species of
rebbachisaurid
Rebbachisauridae is a family of sauropod dinosaurs known from fragmentary fossil remains from the Cretaceous of South America, Africa, North America, Europe and possibly Central Asia.
Taxonomy
In 1990 sauropod specialist Jack McIntosh included t ...
sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their ...
dinosaurs
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
, which lived during the mid-
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
period, about 99.6 to 97 million years ago, in the
Cenomanian, in what is now
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the souther ...
(northwestern
Patagonia
Patagonia () refers to a geographical region that encompasses the southern end of South America, governed by Argentina and Chile. The region comprises the southern section of the Andes Mountains with lakes, fjords, temperate rainforests, and ...
).
Description

''Limaysaurus'' was a medium-sized sauropod.
Gregory S. Paul in 2010 estimated its length at fifteen meters (50 ft) and its weight at seven
tonnes
The tonne ( or ; symbol: t) is a unit of mass equal to 1000 kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton to distinguish it from the non-metric units of the short ton ( United States ...
.
[Paul, G.S., 2010, ''The Princeton Field Guide to Dinosaurs'', Princeton University Press, pp. 186 and 208.] The neural spines on its back were very tall. The neural spines of the cervical and dorsal vertebrae are not V-shaped but they have a simple and straight form like an I. Its teeth were curved, unlike those of ''
Diplodocus
''Diplodocus'' (, , or ) was a genus of diplodocid sauropod dinosaurs, whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a neo-Latin term derived from Greek διπ� ...
'' which were pencil-shaped.
Another distinct characteristic of this sauropod is its
phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
relationship to ''
Rebbachisaurus'' from
Morocco
Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria ...
. This discovery supports the theory that there was a
land bridge
In biogeography, a land bridge is an isthmus or wider land connection between otherwise separate areas, over which animals and plants are able to cross and colonize new lands. A land bridge can be created by marine regression, in which sea ...
connecting South America to
Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
100 million years ago. It shared its habitat with ''
Andesaurus
''Andesaurus'' ( ; "Andes lizard") is a genus of basal titanosaurian sauropod dinosaur which existed during the middle of the Cretaceous Period in South America. Like most sauropods, belonging to one of the largest animals ever to walk the Eart ...
'' and ''
Giganotosaurus
''Giganotosaurus'' ( ) is a genus of theropod dinosaur that lived in what is now Argentina, during the early Cenomanian age of the Late Cretaceous period, approximately 99.6 to 95 million years ago. The holotype specimen was discovered in the ...
'', characterized by plains with large and shallow lagoons. The climate was mild and humid. In fossil remains of ''Limaysaurus'',
gastroliths
A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In othe ...
have been found, which were used to help the stomach to grind food.
Discovery
''Limaysaurus'' includes a single
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
, ''Limaysaurus tessonei'', which was originally referred to the genus ''Rebbachisaurus'' as ''Rebbachisaurus tessonei'', an African species, by
Jorge Calvo and
Leonardo Salgado in 1995.
[Calvo, J. O. and Salgado, L. (1995).]
''Rebbachisaurus tessonei'' sp. nov. A new sauropod from the Albian-Cenomanian of Argentina; new evidence on the origin of the Diplodocidae
" ''Gaia'', 11: 13-33. However, a generic separation was proposed by Salgado,
Alberto Garrido,
Sergio Cocca
Sergio may refer to:
* Sergio (given name), for people with the given name Sergio
* Sergio (carbonado), the largest rough diamond ever found
* ''Sergio'' (album), a 1994 album by Sergio Blass
* ''Sergio'' (2009 film), a documentary film
* ''Se ...
and
Juan Cocca, and the genus ''Limaysaurus'' was named in 2004. The generic name is derived from
Río Limay
The Limay River is an important river in the northwestern Argentine Patagonia (the region of Comahue). It originates at the eastern end of the Nahuel Huapi Lake and flows in a meandering path for about , collecting the waters of several tri ...
which borders the region and from the
specific name Specific name may refer to:
* in Database management systems, a system-assigned name that is unique within a particular database
In taxonomy, either of these two meanings, each with its own set of rules:
* Specific name (botany), the two-part (bino ...
, ''tessonei'', in honor of Lieto Tessone, who found the first and most complete
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
.
Their discovery shed some light on the distribution of
Gondwanan
Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final stages ...
dinosaurs in the mid-Cretaceous period. Several specimens of ''Limaysaurus'' are known, one of which (the holotype) is 80% complete, being one of the most complete Cretaceous sauropods ever found in South America.
The holotype specimen, MUCPv-205, a partial skeleton including the back of the skull, was found in 1988 by Lieto Francisco Tessone, who is honoured in the specific name. It was afterwards collected, together with MUCPv-206, a second fragmentary skeleton, by
José Bonaparte. These two adult specimens likely hailed from the top of the
Candeleros Formation
The Candeleros Formation (formerly known as the Candeleros Member of the "Río Limay Formation") is a geologic formation that crops out in the Río Negro, Neuquén, and Mendoza provinces of northern Patagonia, Argentina. It is the oldest formati ...
, which is dated to the early Cenomanian age. An additional smaller specimen, MUCPv-153, was found nearby, although it belonged to the base of the
Huincul Formation, from the late Cenomanian.
[
''Limaysaurus'' was found 15 km (9 mi) southwest of Villa El Chocón, Picún Leufú Department, Neuquén Province, Patagonia, ]Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
. The sediments belong to the Río Limay Subgroup, in Lohan Cura Formation, at Cerro Aguada del León
Cerro is Spanish for "hill" or "mountain".
Toponyms
;Bolivia:
* Cerro Rico, the "Rich Mountain" containing silver ore near Potosi, Bolivia
;Brazil:
*Cerro Branco, a municipality of Rio Grande do Sul
*Cerro Grande, Rio Grande do Sul, a municipa ...
. These beds appear to date from the Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
-Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± ...
interval,[ although these fossils were later named '']Comahuesaurus
''Comahuesaurus'' (meaning "Comahue lizard") is a genus of sauropod dinosaur of the family Rebbachisauridae. It was found in the Lohan Cura Formation, in Argentina and lived during the Early Cretaceous, Aptian to Albian. The type species is ''C. ...
''.
Classification
The describing authors in 2004 assigned ''Limaysaurus'' to the family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Rebbachisauridae. The rebbachisaurids are a basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
clade within the Diplodocimorpha
Diplodocoidea is a superfamily of sauropod dinosaurs, which included some of the longest animals of all time, including slender giants like ''Supersaurus'', ''Diplodocus'', ''Apatosaurus'', and ''Amphicoelias''. Most had very long necks and long, ...
, and their remains have been found in Cretaceous-age rocks
In geology, rock (or stone) is any naturally occurring solid mass or aggregate of minerals or mineraloid matter. It is categorized by the minerals included, its chemical composition, and the way in which it is formed. Rocks form the Earth's ...
in Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
, South America, and Africa. A cladistic
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived ch ...
analysis of macronarian sauropods (Salgado et al., 2004) demonstrated close affinities between ''Limaysaurus'', the African genera ''Rebbachisaurus'' and '' Nigersaurus'', and the South American genera '' Rayososaurus'' and ''Cathartesaura
''Cathartesaura'' is a genus of rebbachisaurid sauropod dinosaur hailing from the Late Cretaceous strata of the Huincul Formation, at the "La Buitrera" locality, in the Neuquén Basin of Río Negro Province, Argentina. The fossil remains, describ ...
''. [Salgado, L., Garrido, A., Cocca, S. E., and Cocca, J. R. (2004). "Lower Cretaceous rebbachisaurid sauropods from Cerro Aguada Del León, Neuquén Province, northwestern Patagonia, Argentina." ''Journal of Vertebrate Paleontology'', 24(4): 903-912, December 2004.]
References
External links
Limaysaurus (Diplodocoidea) in The Thescelsossaurus
{{Taxonbar, from=Q137252
Rebbachisaurids
Cenomanian life
Early Cretaceous dinosaurs of South America
Cretaceous Argentina
Candeleros Formation
Huincul Formation
Fossils of Argentina
Fossil taxa described in 2004