Leyesaurus Skull In Dorsal View
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''Leyesaurus'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
massospondylid Massospondylidae is a family (biology), family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and AntarcticaHellert, Spencer M. "A New Basal Sauropodomorph from The Early Jurassic Hanson Formation of Antarc ...
sauropodomorph Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lo ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
known from the San Juan Province, northwestern
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
.


Description

''Leyesaurus'' is known from the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
PVSJ 706, a nearly complete
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
with articulated
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
and some
postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...
l remains (
vertebral column The spinal column, also known as the vertebral column, spine or backbone, is the core part of the axial skeleton in vertebrates. The vertebral column is the defining and eponymous characteristic of the vertebrate. The spinal column is a segmente ...
,
scapular A scapular () is a Western Christian garment suspended from the shoulders. There are two types of scapulars, the monastic and devotional scapular; both forms may simply be referred to as "scapular". As an object of popular piety, a scapular ...
and
pelvic girdle The hip bone (os coxae, innominate bone, pelvic bone or coxal bone) is a large flat bone, constricted in the center and expanded above and below. In some vertebrates (including humans before puberty) it is composed of three parts: the Ilium (bone) ...
s and
hindlimb A hindlimb or back limb is one of the paired articulated appendages ( limbs) attached on the caudal ( posterior) end of a terrestrial tetrapod vertebrate's torso.http://www.merriam-webster.com/medical/hind%20limb, Merriam Webster Dictionary-H ...
). The skull has a length of 18
centimeter upright=1.35, Different lengths as in respect to the electromagnetic spectrum, measured by the metre and its derived scales. The microwave is in-between 1 meter to 1 millimeter. A centimetre (International spelling) or centimeter (American ...
s, and ''Leyesaurus'' has been estimated to have been about in length. It was collected from the uppermost part of the Quebrada del Barro Formation of the Marayes-El Carrizal Basin, dating to the
Lower Jurassic The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic–Jurassic extinction event, 201.3 Ma&nb ...
(based on the similarities between ''Leyesaurus'' and Lower Jurassic sauropodomorphs, like ''
Massospondylus ''Massospondylus'' ( ) is a genus of sauropodomorph dinosaur from the Early Jurassic of southern Africa. It was described by Sir Richard Owen in 1854 from remains discovered in South Africa, and is thus one of the first dinosaurs to have been ...
'' and '' Adeopapposaurus''). ''Leyesaurus'' was found near the locality Balde de Leyes, in the Caucete Department of San Juan Province. Within
Massospondylidae Massospondylidae is a family (biology), family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and AntarcticaHellert, Spencer M. "A New Basal Sauropodomorph from The Early Jurassic Hanson Formation of Antarc ...
, ''Leyesaurus'' was found to be most closely related to '' Adeopapposaurus''.


Etymology

''Leyesaurus'' was first named by Cecilia Apaldetti, Ricardo N. Martinez, Oscar A. Alcober and Diego Pol in
2011 The year marked the start of a Arab Spring, series of protests and revolutions throughout the Arab world advocating for democracy, reform, and economic recovery, later leading to the depositions of world leaders in Tunisia, Egypt, and Yemen ...
and the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Leyesaurus marayensis''. The generic name honors the Leyes family, inhabitants of the small town Balde de Leyes, who discovered the holotype and notified the paleontologists of the San Juan Museum, and ''saurus'',
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
for "lizard". The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
refers to the Marayes-El Carrizal Basin, where the holotype was discovered.


Phylogeny

The following cladogram shows the position of ''Leyesaurus'' within Massopoda, according to Oliver W. M. Rauhut and colleagues, 2020:


References

{{Taxonbar, from=Q1656645 Massospondylidae Dinosaur genera Pliensbachian dinosaurs Fossil taxa described in 2011 Dinosaurs of Argentina