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The levator ani is a broad, thin
muscle Muscle is a soft tissue, one of the four basic types of animal tissue. There are three types of muscle tissue in vertebrates: skeletal muscle, cardiac muscle, and smooth muscle. Muscle tissue gives skeletal muscles the ability to muscle contra ...
group, situated on either side of the
pelvis The pelvis (: pelves or pelvises) is the lower part of an Anatomy, anatomical Trunk (anatomy), trunk, between the human abdomen, abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also c ...
. It is formed from three muscle components: the pubococcygeus, the iliococcygeus, and the puborectalis. It is attached to the inner surface of each side of the
lesser pelvis The pelvic cavity is a body cavity that is bounded by the bones of the pelvis. Its oblique roof is the pelvic inlet (the superior opening of the pelvis). Its lower boundary is the pelvic floor. The pelvic cavity primarily contains the reproduc ...
, and these unite to form the greater part of the pelvic floor. The coccygeus muscle completes the pelvic floor, which is also called the ''pelvic diaphragm''. It supports the
viscera In a multicellular organism, an organ is a collection of tissues joined in a structural unit to serve a common function. In the hierarchy of life, an organ lies between tissue and an organ system. Tissues are formed from same type cells to a ...
in the pelvic cavity, and surrounds the various structures that pass through it. The levator ani is the main pelvic floor muscle and contracts rhythmically during female orgasm, and painfully during vaginismus.


Structure

The levator ani is made up of 3 parts: * Iliococcygeus muscle * Pubococcygeus muscle * Puborectalis muscle The iliococcygeus arises from the inner side of the ischium (the lower and back part of the
hip bone The hip bone (os coxae, innominate bone, pelvic bone or coxal bone) is a large flat bone, constricted in the center and expanded above and below. In some vertebrates (including humans before puberty) it is composed of three parts: the Ilium (bone) ...
) and from the posterior part of the tendinous arch of the obturator fascia, and is attached to the coccyx and anococcygeal body; it is usually thin, and may be absent, or be largely replaced by fibrous tissue. An accessory slip at its posterior part is sometimes named the ''iliosacralis''. The pubococcygeus muscle has medial fibres forming the pubovaginalis in the female, and the puboprostaticus in the male.


Origin and insertion

The levator ani arises, in front, from the posterior surface of the superior pubic ramus lateral to the symphysis; behind, from the inner surface of the spine of the ischium; and between these two points, from the obturator fascia. Posteriorly, this fascial origin corresponds, more or less closely, with the tendinous arch of the pelvic fascia, but in front, the muscle arises from the fascia at a varying distance above the arch, in some cases reaching nearly as high as the canal for the obturator vessels and nerve. The fibers pass downward and backward to the middle line of the floor of the pelvis; the most posterior are inserted into the side of the last two segments of the coccyx; those placed more anteriorly unite with the muscle of the opposite side, in a median fibrous ridge called the anococcygeal body or raphe, which extends between the coccyx and the margin of the anus. The middle fibers are inserted into the side of the
rectum The rectum (: rectums or recta) is the final straight portion of the large intestine in humans and some other mammals, and the gut in others. Before expulsion through the anus or cloaca, the rectum stores the feces temporarily. The adult ...
, blending with the fibers of the sphincter muscles; lastly in the male, the anterior fibers descend upon the side of the prostate to unite beneath it with the muscle of the opposite side, joining with the fibers of the external anal sphincter and transverse perineal muscles, at the central tendinous point of the
perineum The perineum (: perineums or perinea) in placentalia, placental mammals is the space between the anus and the genitals. The human perineum is between the anus and scrotum in the male or between the anus and vulva in the female. The perineum is ...
. The anterior portion is occasionally separated from the rest of the muscle by connective tissue. From this circumstance, as well as from its peculiar relation with the
prostate The prostate is an male accessory gland, accessory gland of the male reproductive system and a muscle-driven mechanical switch between urination and ejaculation. It is found in all male mammals. It differs between species anatomically, chemica ...
, which it supports as in a sling, it has been described as a distinct muscle, under the name of ''levator prostatæ''. In the female, the anterior fibers of the levator ani descend upon the side of the
vagina In mammals and other animals, the vagina (: vaginas or vaginae) is the elastic, muscular sex organ, reproductive organ of the female genital tract. In humans, it extends from the vulval vestibule to the cervix (neck of the uterus). The #Vag ...
.


Innervation

The levator ani muscles are mostly innervated by the pudendal nerve, perineal nerve and acting together.


Variation

In addition, sacral spinal nerves ( S3, S4) innervate the muscles directly as well (in ~70% of people). Sometimes (in ~40% of people) the inferior rectal nerve innervates the levator ani muscles independently of the pudendal nerve.


Pubococcygeus muscle

The pubococcygeus muscle or PC muscle is a hammock-like muscle, found in both sexes, that stretches from the pubic bone to the coccyx (tail bone) forming the
floor A floor is the bottom surface of a room or vehicle. Floors vary from wikt:hovel, simple dirt in a cave to many layered surfaces made with modern technology. Floors may be stone, wood, bamboo, metal or any other material that can support the ex ...
of the pelvic cavity and supporting the pelvic organs.


Structure

The pubococcygeus arises from the back of the pubis and from the anterior part of the obturator fascia, and is directed backward almost horizontally along the side of the
anal canal The anal canal is the part that connects the rectum to the anus, located below the level of the pelvic diaphragm. It is located within the anal triangle of the perineum, between the right and left ischioanal fossa. As the final functional s ...
toward the coccyx and sacrum, to which it finds attachment. Between the termination of the vertebral column and the anus, the two pubococcygeus muscles come together and form a thick, fibromuscular layer lying on the raphe (ridge) or ( anococcygeal body) formed by the iliococcygei. The greater part of this muscle is inserted into the coccyx and into the last one or two pieces of the sacrum.


Variation

This insertion into the vertebral column is, however, not accepted by all observers.


Function

The pubococcygeus muscle controls urine flow and contracts during orgasm as well as assisting in male
ejaculation Ejaculation is the discharge of semen (the ''ejaculate''; normally containing sperm) from the penis through the urethra. It is the final stage and natural objective of male sexual stimulation, and an essential component of natural conception. ...
. It also aids in
childbirth Childbirth, also known as labour, parturition and delivery, is the completion of pregnancy, where one or more Fetus, fetuses exits the Womb, internal environment of the mother via vaginal delivery or caesarean section and becomes a newborn to ...
as well as core stability. A strong pubococcygeus muscle has also been linked to a reduction in urinary incontinence and proper positioning of the baby's head during childbirth.


Kegel exercises

The Kegel exercises are a series of voluntary contractions of all the perineal muscles. Such movement is done in an effort to strengthen all the striated muscles in the perineum's area. They are often referred to simply as "kegels", named after their founder, Dr. Arnold Kegel. These exercises also serve to contract, among others, the
ischiocavernosus The ischiocavernosus muscle (erectores penis ''or'' erector clitoridis in older texts) is a muscle just below the surface of the perineum, present in both men and women. Structure It arises by tendinous and fleshy fibers from the inner surface of ...
, bulbospongiosus, and cremaster muscle in men, as voluntary contraction of the pubococcygeus muscle also engages the cremasteric reflex, which lifts the testicles up, although this does not occur in all men. Kegel exercises have been prescribed to ameliorate
erectile dysfunction Erectile dysfunction (ED), also referred to as impotence, is a form of sexual dysfunction in males characterized by the persistent or recurring inability to achieve or maintain a Human penis, penile erection with sufficient rigidity and durat ...
due to venous leakage and to help men control
premature ejaculation Premature ejaculation (PE) is a male sexual dysfunction that occurs when a male Ejaculation, expels semen (and most likely experiences orgasm) soon after beginning sexual activity, and with minimal penile stimulation. It has also been called ''e ...
and to treat urinary incontinence in both sexes.


Puborectalis muscle

The fibers that form a sling looping around the
rectum The rectum (: rectums or recta) is the final straight portion of the large intestine in humans and some other mammals, and the gut in others. Before expulsion through the anus or cloaca, the rectum stores the feces temporarily. The adult ...
are named puborectalis (puboanalis). They arise from the lower part of the pubic symphysis, and from the superior fascia of the urogenital diaphragm. The origin of the puborectalis fibers is at the posterior surface of the pubis while their insertion is at the midline sling posterior to the rectum. The muscle band is innervated by perineal branches of the S3 and S4 nerve roots. They meet with the corresponding fibers of the opposite side around the lower part of the rectum, and form for it a strong sling. Relaxation increases the angle between rectum and anus, allowing defecation in conjunction with relaxation of the internal and external anal sphincters. Levator ani relaxation and rectal emptying is facilitated by anorectal straightening during squatting.


Function

The levator ani muscles are responsible for "wagging" the tail in
tail The tail is the elongated section at the rear end of a bilaterian animal's body; in general, the term refers to a distinct, flexible appendage extending backwards from the midline of the torso. In vertebrate animals that evolution, evolved to los ...
ed
quadruped Quadrupedalism is a form of locomotion in which animals have four legs that are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four legs is said to be a quadruped (fr ...
s. These muscles are not as strong in the human, as tail-wagging is more demanding than the support function that the muscles serve in humans.


Clinical significance


Levator ani syndrome

Levator ani syndrome is episodic rectal pain caused by spasm of the levator ani muscle.Levator Syndrome, by Parswa Ansari, MD 7/2014, Merck Manuals


Levator ani avulsion


Additional images

File:Gray236.png, Right hip bone. Internal surface. File:Gray402.png, Coronal section of pelvis, showing arrangement of fasciæ. Viewed from behind. File:1116 Muscle of the Male Perineum.png, Muscles of male perineum. File:Levator ani.png File:Gray539.png, The arteries of the pelvis. File:Gray837.png, Sacral plexus of the right side. File:Gray1076.png, Iliac colon, sigmoid or pelvic colon, and rectum seen from the front, after removal of pubic bones and bladder. File:Gray1077.png, The posterior aspect of the rectum exposed by removing the lower part of the sacrum and the coccyx. File:Gray1136.png, Male pelvic organs seen from right side. File:Human anus-en.svg, Anatomy of the human anus.


See also

* Bulbospongiosus muscle * Coccygectomy


References


External links

* —"Muscles of the female superficial perineal pouch." * —"Muscles of the male superficial perineal pouch." * —"Muscles of the Pelvic Diaphragm" * * * * * () * (, )
Merck Manual article on levator ani syndrome
{{DEFAULTSORT:Levator Ani Muscles of the torso Pelvis Sexual anatomy