The shagreen ray (''Leucoraja fullonica''), also known as shagreen skate or fuller's ray, is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of skate in the family Rajidae. This ray is found in the eastern Atlantic Ocean, from

Murmansk Murmansk () is a port city and the administrative center of Murmansk Oblast in the far Far North (Russia), northwest part of Russia. It is the world's largest city north of the Arctic Circle and sits on both slopes and banks of a modest fjord, Ko ...

, Russia through Norway, southern Iceland, the

Faroe Islands The Faroe Islands ( ) (alt. the Faroes) are an archipelago in the North Atlantic Ocean and an autonomous territory of the Danish Realm, Kingdom of Denmark. Located between Iceland, Norway, and the United Kingdom, the islands have a populat ...

, the

Celtic Sea The Celtic Sea is the area of the Atlantic Ocean off the southern coast of Ireland bounded to the north by St George's Channel, Saint George's Channel; other limits include the Bristol Channel, the English Channel, and the Bay of Biscay, as wel ...

, the northern

North Sea The North Sea lies between Great Britain, Denmark, Norway, Germany, the Netherlands, Belgium, and France. A sea on the European continental shelf, it connects to the Atlantic Ocean through the English Channel in the south and the Norwegian Se ...

and

Skagerrak The Skagerrak (; , , ) is a strait running between the North Jutlandic Island of Denmark, the east coast of Norway and the west coast of Sweden, connecting the North Sea and the Kattegat sea. The Skagerrak contains some of the busiest shipping ...

, to western Morocco and the

Madeira Madeira ( ; ), officially the Autonomous Region of Madeira (), is an autonomous Regions of Portugal, autonomous region of Portugal. It is an archipelago situated in the North Atlantic Ocean, in the region of Macaronesia, just under north of ...

archipelago, but not in the shallow waters off

England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...

and

Wales Wales ( ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by the Irish Sea to the north and west, England to the England–Wales border, east, the Bristol Channel to the south, and the Celtic ...

. It is also recorded sporadically in the

Mediterranean Sea The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern Eur ...

.Guide of Mediterranean Skates and Rays (''Leucoraja fullonica''). Oct. 2022. Mendez L., Bacquet A. and F. Briand. http://www.ciesm.org/Guide/skatesandrays/leucoraja-fullonica

Shagreen Shagreen is a type of rawhide consisting of rough untanned skin, historically from a horse's or onager's back, or from shark or ray. Etymology The word derives from the French ' and is related to Italian and Venetian ''sagrin'', deriv ...

is a form of roughened untanned leather, typically dyed green and commonly made of the skins of sharks and rays. The

common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...

"fuller's ray" comes from an instrument used by fullers to smooth

cloth Textile is an umbrella term that includes various fiber-based materials, including fibers, yarns, filaments, threads, and different types of fabric. At first, the word "textiles" only referred to woven fabrics. However, weaving is n ...

, referring to its rough and spiny back. The

pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...

disc of the shagreen ray is somewhat wider than it is long. The forward margin

concave Concave or concavity may refer to: Science and technology * Concave lens * Concave mirror Mathematics * Concave function, the negative of a convex function * Concave polygon A simple polygon that is not convex is called concave, non-convex or ...

and undulated, and about one-third longer than the rear margin. The

snout A snout is the protruding portion of an animal's face, consisting of its nose, mouth, and jaw. In many animals, the structure is called a muzzle, Rostrum (anatomy), rostrum, beak or proboscis. The wet furless surface around the nostrils of the n ...

is cone-shaped and pointed. The

tail The tail is the elongated section at the rear end of a bilaterian animal's body; in general, the term refers to a distinct, flexible appendage extending backwards from the midline of the torso. In vertebrate animals that evolution, evolved to los ...

is shorter than the disc, with two small, equally sized

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

s and a rudimentary

caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...

. The

eye An eye is a sensory organ that allows an organism to perceive visual information. It detects light and converts it into electro-chemical impulses in neurons (neurones). It is part of an organism's visual system. In higher organisms, the ey ...

s and

mouth A mouth also referred to as the oral is the body orifice through which many animals ingest food and animal communication#Auditory, vocalize. The body cavity immediately behind the mouth opening, known as the oral cavity (or in Latin), is also t ...

are large; the

teeth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...

are pointed and number about 64 in the upper jaw and 56 in the

lower Lower may refer to: * ''Lower'' (album), 2025 album by Benjamin Booker *Lower (surname) *Lower Township, New Jersey *Lower Receiver (firearms) *Lower Wick Lower Wick is a small hamlet located in the county of Gloucestershire, England. It is sit ...

. The upper surface is rough in texture, with patches of large spines on the snout, around the eyes, and on the shoulder. There are two rows of about 50 spines each running from the center of the back to the first dorsal fin, and no interdorsal spine. The underside is smooth except for the front edge of the disc, base of the tail, and tail. The coloration is a uniform gray or brown with smaller darker dots above, and white below. The maximum size is to 120 cm long. This species occurs in relatively cold water on the upper

continental slope A continental margin is the outer edge of continental crust abutting oceanic crust under coastal waters. It is one of the three major zones of the ocean floor, the other two being deep-ocean basins and mid-ocean ridges. The continental margi ...

s at a depth of 30–550 m. It is most common at depths of 200 m, but is found deeper in southern areas. It favors

sand Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is usually defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural ...

y and possibly also rocky

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s. The shagreen ray feeds on a variety of

benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". ...

animal Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, ...

s, mainly

fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...

es but also

crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...

s. Like other skates reproduction is

oviparous Oviparous animals are animals that reproduce by depositing fertilized zygotes outside the body (i.e., by laying or spawning) in metabolically independent incubation organs known as eggs, which nurture the embryo into moving offsprings kno ...

; the

egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...

capsules are amber in color and oblong-shaped, with long, stiff horns at each corner. One pair of horns are longer than the capsule. The capsules measure 7.5-9.9 cm long and 4.5-4.7 cm wide. Historically this ray was considered of "inferior" quality for consumption. It is caught commercially by longlines and trawls.

References

{{Taxonbar, from=Q383482 shagreen ray Fish of the North Sea Marine fish of Europe shagreen ray shagreen ray