''Leucocarbo'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of birds in the family Phalacrocoracidae. Several species within the genus are collectively known as blue-eyed shags. This is a group of closely related

cormorant Phalacrocoracidae is a family of approximately 40 species of aquatic birds commonly known as cormorants and shags. Several different classifications of the family have been proposed, but in 2021 the International Ornithologists' Union (IOU) ado ...

taxa. Many have a blue, purple or red ring around the eye (not a blue iris); other shared features are white underparts (at least in some individuals) and pink feet. They are found around the colder parts of the Southern Hemisphere, especially near southern

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...

, and

New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...

. Many are

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to remote islands. Determining which types are species and which are subspecies of what larger species is problematic; various recent authorities have recognized from 8 to 14 species and have placed them in a variety of genera. The common names are even more confusing, "like myriad footprints criss-crossing in the snow and about as easy to disentangle." Only one common name is given for most species here.

Taxonomy

The genus ''Leucocarbo'' was introduced in 1856 by the French naturalist

Charles Lucien Bonaparte Charles Lucien Jules Laurent Bonaparte, 2nd Prince of Canino and Musignano (24 May 1803 – 29 July 1857) was a French naturalist and ornithology, ornithologist, and a nephew of Napoleon. Lucien and his wife had twelve children, including Cardinal ...

. He did not specify a

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

but this was designated as the guanay cormorant by William Ogilvie-Grant in 1898. The name ''Leucocarbo'' combines the

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

''leukos'' meaning "white" with the genus name ''Carbo'' introduced by

Bernard Germain de Lacépède Bernard-Germain-Étienne de La Ville-sur-Illon, comte de Lacépède or La Cépède (; 26 December 17566 October 1825) was a French natural history, naturalist and an active freemason. He is known for his contribution to the Comte de Buffon's g ...

in 1799. A

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

study published in 2014 found that ''Leucocarbo'' is

sister A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to ref ...

to the American cormorants in the genus ''

Nannopterum ''Nannopterum'' is a genus of cormorant comprising three species. They are found throughout the Americas, hence the common name American cormorants. These species were formerly classified in the genus ''Phalacrocorax''. A molecular phylogenetic ...

''; the genera split between 6.7 - 8.0 million years ago. The genus contains 15 species:

References

{{DEFAULTSORT:Blue-Eyed Shag Leucocarbo Taxa named by Charles Lucien Bonaparte