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The flat pebblesnail (''Lepyrium showalteri'') is a
species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of
freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
, an aquatic gastropod
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is es ...
in the family
Lithoglyphidae Lithoglyphidae is a family of small freshwater snails with gills and an operculum, aquatic gastropod mollusks. This family is in the superfamily Truncatelloidea and in the clade Littorinimorpha (according to the taxonomy of the Gastropoda by ...
. This species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
to the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
. This snail has been listed as endangered on the
United States Fish and Wildlife Service list of endangered species This is a list of the bird and mammal species and subspecies described as endangered by the United States Fish and Wildlife Service. It contains species and subspecies not only in the U.S. and its territories, but also those only found in other par ...
since October 28, 1998. ''Lepyrium showalteri'' is the only species in the genus ''Lepyrium''. ''Lepyrium showalteri'' is the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
of the genus ''Lepyrium''. Lepyriidae is a synonym of Lithoglyphinae.


Description

The flat pebblesnail is a small snail in the family Lithoglyphidae; however, the species has a large and distinctive-looking
shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses ** Thin-shell structure Science Biology * Seashell, a hard o ...
. This snail's shell is also distinguished by its depressed
spire A spire is a tall, slender, pointed structure on top of a roof of a building or tower, especially at the summit of church steeples. A spire may have a square, circular, or polygonal plan, with a roughly conical or pyramidal shape. Spires are ...
and expanded, flattened body whorl. The shells are ovate in outline, flattened, and grow to 3.5 to 4.4 mm (0.1 to 0.2 in) high and 4 to 5 mm (0.2 in) wide. The
umbilical Umbilical may refer to: *Umbilical cable *Umbilical cord *Umbilical fold *Umbilical hernia *Umbilical notch *Umbilical vessels **Umbilical artery **Umbilical vein *Umbilical zone *The Umbilical Brothers, two Australian comedic performers, David a ...
area is imperforate (no opening), and there are 2 to 3 whorls which rapidly expand. The anatomy of this species was described in detail by Thompson in 1984.Thompson F. G. (1984). "North American freshwater snail general of the hydrobiid subfamily Lithoglyphinae". ''
Malacologia ''Malacologia'' is a peer-reviewed scientific journal in the field of malacology, the study of mollusks. The journal publishes articles in the fields of molluscan systematics, ecology, population ecology, genetics, molecular genetics, evolution, a ...
'' 25(1)
109
141.


Original description

Species ''Lepyrium showalteri'' was originally described as ''Neritina showalteri'' by
Isaac Lea Isaac; grc, Ἰσαάκ, Isaák; ar, إسحٰق/إسحاق, Isḥāq; am, ይስሐቅ is one of the three patriarchs of the Israelites and an important figure in the Abrahamic religions, including Judaism, Christianity, and Islam. He was t ...
in 1861. Lea's original text (the
type description A species description is a formal description of a newly discovered species, usually in the form of a scientific paper. Its purpose is to give a clear description of a new species of organism and explain how it differs from species that have ...
) reads as follows:


Distribution

This species is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
to
Alabama (We dare defend our rights) , anthem = " Alabama" , image_map = Alabama in United States.svg , seat = Montgomery , LargestCity = Huntsville , LargestCounty = Baldwin County , LargestMetro = Greater Birmingham , area_total_km2 = 135,7 ...
in the United States. It was historically known from: * the mainstem
Coosa River The Coosa River is a tributary of the Alabama River in the U.S. states of Alabama and Georgia. The river is about long.U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed April 27, 2011 ...
in Shelby County and
Talladega County, Alabama Talladega County (pronounced Talla-dig-a) is a county located in the east central portion of the U.S. state of Alabama."ACES Winston County Office" (links/history), Alabama Cooperative Extension System (ACES), 2007, webpageACES-Talladega As of ...
; * the
Cahaba River The Cahaba River is the longest substantially free-flowing river in Alabama and is among the most scenic and biologically diverse rivers in the United States. It is a major tributary of the Alabama River and part of the larger Mobile River basin. ...
in
Bibb Bibb may refer to: Places in the United States * Bibb County, Alabama * Fort Bibb, Alabama, constructed in 1818 * Bibb City, Georgia, in Muscogee County * Bibb County, Georgia People * Bibb (surname) * Bibb (given name) Ships * , a Revenue Mari ...
and
Dallas County, Alabama Dallas County is a county located in the central part of the U.S. state of Alabama. As of the 2020 census, its population was 38,462. The county seat is Selma. Its name is in honor of United States Secretary of the Treasury Alexander J. Dalla ...
; * and the
Little Cahaba River Little is a synonym for small size and may refer to: Arts and entertainment * ''Little'' (album), 1990 debut album of Vic Chesnutt * ''Little'' (film), 2019 American comedy film * The Littles, a series of children's novels by American author John ...
in Bibb County, Alabama. The flat pebblesnail has not been found in the Coosa River portion of its range since the construction of the
Lay Dam Lay Dam is a hydroelectric power dam on the Coosa River in Chilton County and Coosa County, near Clanton, Alabama.Tharpe, Bill"Lay Dam and Lake" encyclopediaofalabama.org, January 7, 2009, updated October 5, 2010. The concrete run-of-the-river ...
and Logan Martin Dam, and recent survey efforts have failed to locate any surviving populations outside of the Cahaba River drainage.Bogan A. E. & Pierson J. M. (1993). ''Survey of the aquatic gastropods of the Cahaba River Basin, Alabama: 1992''. Alabama Natural Heritage Program. Contract Number 1922. The flat pebblesnail is currently known from one site on the
Little Cahaba River Little is a synonym for small size and may refer to: Arts and entertainment * ''Little'' (album), 1990 debut album of Vic Chesnutt * ''Little'' (film), 2019 American comedy film * The Littles, a series of children's novels by American author John ...
, Bibb County, and from a single shoal series on the
Cahaba River The Cahaba River is the longest substantially free-flowing river in Alabama and is among the most scenic and biologically diverse rivers in the United States. It is a major tributary of the Alabama River and part of the larger Mobile River basin. ...
above the Fall Line,
Shelby County, Alabama Shelby County is located in the central portion of the U.S. state of Alabama. As of the 2020 census the population was 223,024. The county seat is Columbiana. The largest city is Alabaster. The county is named in honor of Isaac Shelby, Govern ...
. File:Map of Alabama highlighting Bibb County.svg,
Bibb County, Alabama Bibb County is a county in the central portion of the U.S. state of Alabama. The county is included in the ARC's definition of Appalachia. As of the 24th decennial 2020 census, its population was 22,293. The county seat is Centreville. The ...
(marked red) where the species is found in the
Little Cahaba River Little is a synonym for small size and may refer to: Arts and entertainment * ''Little'' (album), 1990 debut album of Vic Chesnutt * ''Little'' (film), 2019 American comedy film * The Littles, a series of children's novels by American author John ...
. File:Map of Alabama highlighting Shelby County.svg,
Shelby County, Alabama Shelby County is located in the central portion of the U.S. state of Alabama. As of the 2020 census the population was 223,024. The county seat is Columbiana. The largest city is Alabaster. The county is named in honor of Isaac Shelby, Govern ...
(marked red) where the species is known from the
Cahaba River The Cahaba River is the longest substantially free-flowing river in Alabama and is among the most scenic and biologically diverse rivers in the United States. It is a major tributary of the Alabama River and part of the larger Mobile River basin. ...
.


Ecology

Little is known of the natural history of this species. The flat pebblesnail is found attached to clean, smooth stones in rapid currents of river shoals. The eggs are laid in capsules on hard surfaces. The life span appears to be 1 year.Johnson P. pers. comm. (2005). In: U.S. Fish and Wildlife Service (2005)
''Recovery Plan for 6 Mobile River Basin Aquatic Snails''
U.S. Fish and Wildlife Service, Jackson, Mississippi. 46 pp., page 5-6.


References

This article incorporates public domain text from the reference and a
public domain The public domain (PD) consists of all the creative work to which no exclusive intellectual property rights apply. Those rights may have expired, been forfeited, expressly waived, or may be inapplicable. Because those rights have expired, ...
work of the United States Government from the reference
U.S. Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
(2005)
''Recovery Plan for 6 Mobile River Basin Aquatic Snails''
U.S. Fish and Wildlife Service, Jackson, Mississippi. 46 pp., page 5-6.


External links

* Tryon G. W. (1888-1889) ''
Manual of Conchology George Washington Tryon Jr. (20 May 1838 – 5 February 1888) was an American malacologist who worked at the Academy of Natural Sciences in Philadelphia. Biography George Washington Tryon was the son of Edward K. Tryon and Adeline Savidt. ...
, structural and systematic, with illustrations of the species''. 10
53plate 17
figure 81-82. {{Taxonbar, from=Q3173245 Gastropods described in 1861 Endemic fauna of Alabama Lithoglyphidae Taxonomy articles created by Polbot ESA endangered species