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The lemon (''Citrus'' × ''limon'') is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of small
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
tree in the ''
Citrus ''Citrus'' is a genus of flowering trees and shrubs in the family Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, mandarins, lemons, grapefruits, pomelos, and limes. ''Citrus'' is nativ ...
'' genus of the
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
family
Rutaceae The Rutaceae () is a family (biology), family, commonly known as the rueRUTACEAE
in Bo ...
. A true lemon is a hybrid of the citron and the bitter orange. Its origins are uncertain, but some evidence suggests lemons originated during the 1st millennium BC in what is now northeastern
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
. Some other citrus fruits are called ''lemon''. The yellow
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
of the lemon tree is used throughout the world, primarily for its juice. The pulp and rind are used in cooking and baking. The juice of the lemon is about 5–6% citric acid, giving it a sour taste. This makes it a key ingredient in
drink A drink or beverage is a liquid intended for human consumption. In addition to their basic function of satisfying thirst, drinks play important roles in human culture. Common types of drinks include plain drinking water, milk, juice, smoothie ...
s and foods such as lemonade and lemon meringue pie. In 2022, world production was 22 million
tonne The tonne ( or ; symbol: t) is a unit of mass equal to 1,000  kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton in the United States to distinguish it from the non-metric units of the s ...
s, led by India with 18% of the total.


Description

The lemon tree produces a pointed oval yellow
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
. Botanically this is a hesperidium, a modified berry with a tough, leathery rind. The rind is divided into an outer colored layer or zest, which is aromatic with essential oils, and an inner layer of white spongy pith. Inside are multiple carpels arranged as radial segments. The seeds develop inside the carpels. The space inside each segment is a locule filled with juice vesicles. Lemons contain many phytochemicals, including
polyphenol Polyphenols () are a large family of naturally occurring phenols. They are abundant in plants and structurally diverse. Polyphenols include phenolic acids, flavonoids, tannic acid, and ellagitannin, some of which have been used historically as ...
s,
terpene Terpenes () are a class of natural products consisting of compounds with the formula (C5H8)n for n ≥ 2. Terpenes are major biosynthetic building blocks. Comprising more than 30,000 compounds, these unsaturated hydrocarbons are produced predomi ...
s, and tannins. Their juice contains slightly more citric acid than lime juice (about 47 g/L), nearly twice as much as grapefruit juice, and about five times as much as orange juice.


Origins

The lemon, like many other cultivated ''
Citrus ''Citrus'' is a genus of flowering trees and shrubs in the family Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, mandarins, lemons, grapefruits, pomelos, and limes. ''Citrus'' is nativ ...
'' species, is a hybrid, in its case of the citron and the bitter orange. and Supplement Lemons were most likely first grown in northeast India. The origin of the word ''lemon'' may be Middle Eastern. The word draws from the Old French ''limon'', then Italian ''limone'', from the Arabic ''laymūn'' or ''līmūn'', and from the Persian ''līmūn'', a generic term for
citrus ''Citrus'' is a genus of flowering trees and shrubs in the family Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, mandarins, lemons, grapefruits, pomelos, and limes. ''Citrus'' is nativ ...
fruit, which is a cognate of Sanskrit (''nimbū'', ' lime'). Lemons entered
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
near southern
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
no later than the second century AD, during the time of
Ancient Rome In modern historiography, ancient Rome is the Roman people, Roman civilisation from the founding of Rome, founding of the Italian city of Rome in the 8th century BC to the Fall of the Western Roman Empire, collapse of the Western Roman Em ...
. They were later introduced to
Persia Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
and then to
Iraq Iraq, officially the Republic of Iraq, is a country in West Asia. It is bordered by Saudi Arabia to Iraq–Saudi Arabia border, the south, Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq border, the east, the Persian Gulf and ...
and
Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...
around 700 AD. The lemon was first recorded in literature in a 10th-century
Arabic Arabic (, , or , ) is a Central Semitic languages, Central Semitic language of the Afroasiatic languages, Afroasiatic language family spoken primarily in the Arab world. The International Organization for Standardization (ISO) assigns lang ...
treatise on farming; it was used as an ornamental plant in early Islamic gardens. It was distributed widely throughout the Arab world and the Mediterranean region in the Arab Agricultural Revolution between 1000 and 1150. A section on lemon and lime tree cultivation in Andalusia,
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
, was included in Ibn al-'Awwam's 12th-century agricultural work, ("Book on Agriculture"). The first substantial cultivation of lemons in Europe began in Genoa in the middle of the 15th century. It was introduced to the
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
in 1493, when Christopher Columbus brought lemon seeds to Hispaniola on his voyages. Spanish conquest throughout the
New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...
helped spread lemon seeds, part of the
Columbian exchange The Columbian exchange, also known as the Columbian interchange, was the widespread transfer of plants, animals, and diseases between the New World (the Americas) in the Western Hemisphere, and the Old World (Afro-Eurasia) in the Eastern Hemis ...
of plants between the Old and New Worlds. It was mainly used as an ornamental plant and for medicine. In the 19th century, lemons were increasingly planted in
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
and
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
. In 1747, the English physician James Lind's experiments on seamen suffering from scurvy involved adding lemon juice to their diets, though vitamin C was not yet known as an important dietary ingredient.


Cultivation


Growing and pruning

Lemons need a minimum temperature of around , so they are not hardy year-round in temperate climates, but become hardier as they mature. Citrus require minimal pruning by trimming overcrowded branches, with the tallest branch cut back to encourage bushy growth. Throughout summer, pinching back tips of the most vigorous growth assures more abundant canopy development. As mature plants may produce unwanted, fast-growing shoots (called "water shoots"), these are removed from the main branches at the bottom or middle of the plant. There is reputed merit in the tradition of urinating near a lemon tree. In cultivation in the UK, the cultivars "Meyer" and "Variegata" have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...
(confirmed 2017).


Production

In 2022, world production of lemons (combined with limes for reporting) was 22 million
tonne The tonne ( or ; symbol: t) is a unit of mass equal to 1,000  kilograms. It is a non-SI unit accepted for use with SI. It is also referred to as a metric ton in the United States to distinguish it from the non-metric units of the s ...
s led by India with 18% of the total. Mexico and China were major secondary producers (table).


Varieties

The 'Bonnie Brae' is oblong, smooth, thin-skinned, and seedless. These are mostly grown in San Diego County, US. The 'Eureka' grows year-round and abundantly. This is the common supermarket lemon, also known as "Four Seasons" (''Quatre Saisons'') because of its ability to produce fruit and flowers together throughout the year. This variety is also available as a plant for domestic customers. There is also a pink-fleshed Eureka lemon with a green and yellow variegated outer skin. The Lisbon lemon is very similar to the Eureka and is the other common supermarket lemon. It is smoother than the Eureka, has thinner skin, and has fewer or no seeds. It generally produces more juice than the Eureka. The 'Femminello St. Teresa', or 'Sorrento' originates in Italy. This fruit's zest is high in lemon oils. It is the variety traditionally used in the making of '' limoncello''. The 'Yen Ben' is an Australasian cultivar.


Uses


Nutrition

Lemon is a rich source of vitamin C, providing 64% of the Daily Value in a 100 g reference amount (table). Other essential nutrients are low in content.


Culinary

Lemon juice and rind are used in a wide variety of foods and drinks, the juice for its sour taste, from its content of 5–6% citric acid. The whole lemon is used to make marmalade, lemon curd and lemon liqueurs such as Limoncello. Lemon slices and lemon rind are used as a garnish for food and drinks. Lemon zest, the grated outer rind of the fruit, is used to add flavor to baked goods. The juice is used to make lemonade and some
cocktail A cocktail is a mixed drink, usually alcoholic beverage, alcoholic. Most commonly, a cocktail is a combination of one or more liquor, spirits mixed with other ingredients, such as juices, flavored syrups, tonic water, Shrub (drink), shrubs, and ...
s. It is used in marinades for fish, where its acid neutralizes amines in fish. In meat, the acid partially hydrolyzes tough collagen fibers, tenderizing it. In the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
, lemon juice is frequently added to
pancakes A pancake, also known as a hotcake, griddlecake, or flapjack, is a flat type of batter bread like cake, often thin and round, prepared from a starch-based Batter (cooking), batter that may contain eggs, milk, and butter, and then cooked on a ...
eaten to celebrate Shrove Tuesday. Lemon juice is used as a short-term
preservative A preservative is a substance or a chemical that is added to products such as food products, beverages, pharmaceutical drugs, paints, biological samples, cosmetics, wood, and many other products to prevent decomposition by microbial growth or ...
on certain foods that tend to oxidize and turn brown after being sliced ( enzymatic browning), such as
apple An apple is a round, edible fruit produced by an apple tree (''Malus'' spp.). Fruit trees of the orchard or domestic apple (''Malus domestica''), the most widely grown in the genus, are agriculture, cultivated worldwide. The tree originated ...
s, bananas, and avocados: its acidity suppresses
oxidation Redox ( , , reduction–oxidation or oxidation–reduction) is a type of chemical reaction in which the oxidation states of the reactants change. Oxidation is the loss of electrons or an increase in the oxidation state, while reduction is ...
by polyphenol oxidase enzymes. Lemon peel is used in the manufacture of pectin, a gelling agent and stabilizer in food and other products. In Mediterranean countries including
Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
, lemons are preserved in jars or barrels of salt. The salt penetrates the peel and rind, softening them, and curing them so that they last almost indefinitely. Lemon oil is extracted from oil-containing cells in the skin. A machine breaks up the cells and uses a water spray to flush off the oil. The oil–water mixture is then filtered and separated by centrifugation. The leaves of the lemon tree are used to make a tea and for preparing cooked meats and seafoods. File:Salmon roll sushi with lemon garnish, September 2008.jpg, Lemon as a garnish File:Oysters served on ice, with lemon and parsley.jpg, Oysters with lemon File:Tenglong Chinese Restaurant Lemon Chicken (37577556406).jpg, Lemon chicken File:A Lemon Meringue Tart.jpg, Lemon meringue pie File:Lemonade - 27682817724.jpg, Lemonade File:Limoncello ready for drinking! (5585765455).jpg, Limoncello


Other uses

Lemons were the primary commercial source of citric acid before the development of fermentation-based processes. Lemon oil is used in aromatherapy. Lemon oil aroma does not influence the human immune system, but may contribute to relaxation. An educational science experiment involves attaching electrodes to a lemon and using it as a battery to produce electricity. Although very low power, several lemon batteries can power a small digital watch. Lemon juice forms a simple invisible ink, developed by heat. Lemon juice is sometimes used to increase the blonde color of hair, acting as a natural highlight after the moistened hair is exposed to sunlight. This works because citric acid acts as bleach.


Other citrus called 'lemons'

* Flat lemon, a mandarin hybrid. * Meyer lemon, a cross between a citron and a mandarin/pomelo hybrid distinct from sour or sweet orange, * Ponderosa lemon, more cold-sensitive than true lemons, the fruit are thick-skinned and very large. Genetic analysis showed it to be a complex hybrid of citron and pomelo. * Rough lemon, a citron-mandarin cross, cold-hardy and often used as a citrus rootstock * Sweet lemons or sweet limes, a mixed group including the lumia (pear lemon), limetta, and Palestinian sweet lime. Among them is the Jaffa lemon, a pomelo-citron hybrid. * Volkamer lemon, like the rough lemon, a citron-mandarin cross


In art and culture

Lemons appear in paintings, pop art, and novels. A wall painting in the tomb of Nakht in 15th century BC Egypt depicts a woman in a festival, holding a lemon. In the 17th century, Giovanna Garzoni painted a ''Still Life with Bowl of Citrons'', the fruits still attached to leafy flowering twigs, with a wasp on one of the fruits. The impressionist Edouard Manet depicted a lemon on a pewter plate. In modern art, Arshile Gorky painted ''Still Life with Lemons'' in the 1930s. In
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
, a lemon may be ritually encircled around a person in the belief that it repels negative energies. It is a common practice for Hindu owners of a new car to drive over four lemons, one under each wheel, crushing them during their first drive. This is believed to protect the driver from accidents. Hindu deities are sometimes depicted with lemons in their iconography, representing the attribute of wealth or abundance. In 20th century American self-improvement culture, Dale Carnegie advised readers "If You Have a Lemon, Make a Lemonade", meaning to make the best of what you have. In the 21st century, a defective machine such as a car is called a lemon. File:Tomb of Nakht.jpg, Fresco in the Tomb of Nakht, woman with lemons, 15th century BC File:Jacob_Foppen_van_Es_-_Still_life_with_pitcher.jpg, ''Still life with pitcher'', Jacob Foppens van Es, between 1617 and 1666 File:Giovanna Garzoni (Italian) - Still Life with Bowl of Citrons - Google Art Project.jpg, ''Still Life with Bowl of Citrons'', Giovanna Garzoni, late 1640s File:Édouard Manet - Le citron.jpg, ''Lemon'',
Édouard Manet Édouard Manet (, ; ; 23 January 1832 – 30 April 1883) was a French Modernism, modernist painter. He was one of the first 19th-century artists to paint modern life, as well as a pivotal figure in the transition from Realism (art movement), R ...
, 1880 File:Lemon by Arshile Gorky 1930s.jpg, ''Still Life with Lemons'', Arshile Gorky, early 1930s


See also

* Limey – the original ration for British sailors was lemon juice


References


External links

* * * * (with illustrations) {{Authority control Citrus hybrids Cocktail garnishes Fruit trees Sour fruits