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The Lau event was the last of three relatively minor
mass extinction An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp fall in the diversity and abundance of multicellular organisms. It occ ...
s (the Ireviken,
Mulde The Mulde () is a river in Saxony and Saxony-Anhalt, Germany. It is a left tributary of the Elbe and is long. The river is formed by the confluence, near Colditz, of the Zwickauer Mulde (running through Zwickau) and the Freiberger Mulde (wit ...
, and Lau events) during the
Silurian The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of t ...
period. It had a major effect on the
conodont Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
fauna, but barely scathed the
graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian ( Miaolingian, Wuliuan) through t ...
s, though they suffered an extinction very shortly thereafter termed the Kozlowskii event that some authors have suggested was coeval with the Lau event and only appears asynchronous due to
taphonomic Taphonomy is the study of how organisms decay and become fossilized or preserved in the paleontological record. The term ''taphonomy'' (from Greek , 'burial' and , 'law') was introduced to paleontology in 1940 by Soviet scientist Ivan Efremov ...
reasons. It coincided with a global low point in sea level caused by glacioeustasy and is closely followed by an excursion in
geochemical Geochemistry is the science that uses the tools and principles of chemistry to explain the mechanisms behind major geological systems such as the Earth's crust and its oceans. The realm of geochemistry extends beyond the Earth, encompassing the ...
isotope Isotopes are distinct nuclear species (or ''nuclides'') of the same chemical element. They have the same atomic number (number of protons in their Atomic nucleus, nuclei) and position in the periodic table (and hence belong to the same chemica ...
s in the ensuing late Ludfordian
faunal stage In chronostratigraphy, a stage is a succession of rock strata laid down in a single age on the geologic timescale, which usually represents millions of years of deposition. A given stage of rock and the corresponding age of time will by conven ...
and a change in depositional regime.


Biological impact

The Lau event started at the beginning of the late Ludfordian, a subdivision of the Ludlow stage, about . Its
strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...
are best exposed in
Gotland Gotland (; ; ''Gutland'' in Gutnish), also historically spelled Gottland or Gothland (), is Sweden's largest island. It is also a Provinces of Sweden, province/Counties of Sweden, county (Swedish län), Municipalities of Sweden, municipality, a ...
,
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
, taking its name from the parish of Lau. Its base is set at the first extinction datum, in the Eke beds, and despite a scarcity of data, it is apparent that most major groups suffered an increase in extinction rate during the event; major changes are observed worldwide at correlated rocks, with a "crisis" observed in populations of
conodont Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
s and
graptolite Graptolites are a group of colonial animals, members of the subclass Graptolithina within the class Pterobranchia. These filter-feeding organisms are known chiefly from fossils found from the Middle Cambrian ( Miaolingian, Wuliuan) through t ...
s. More precisely, conodonts suffered in the Lau event, and graptolites in the subsequent isotopic excursion. Local extinctions may have played a role in many places, especially the increasingly enclosed Welsh basin; the event's relatively high severity rating of 6.2 does not change the fact that many life-forms became re-established shortly after the event, presumably surviving in refuge or in environments that have not been preserved in the geological record. Based on its timing, it's possible that this event finished off the palaeoscolecids. Although life persisted after the event, community structures were permanently altered and many lifeforms failed to regain the niches they had occupied before the event.


Isotopic effects

A peak in , accompanied by fluctuations in other isotope concentrations, is often associated with mass extinctions. Some workers have attempted to explain this event in terms of climate or sea level change – perhaps arising due to a build-up of glaciers; however, such factors alone do not appear to be sufficient to explain the events. An alternative hypothesis is that changes in ocean mixing were responsible. An increase in density is required to make water downwell; the cause of this densification may have changed from hypersalinity (due to ice formation and evaporation) to temperature (due to water cooling). A different hypothesis attributes the carbon isotope fluctuations to methanogenesis caused by the increased influx of iron-bearing dust and consequent disruption of limiting nutrient ratios. Loydell suggests many causes of the isotopic excursion, including increased carbon burial, increased carbonate weathering, changes in atmospheric and oceanic interactions, changes in primary production, and changes in humidity or aridity. He uses a correlation between the events and glacially induced global sea level change to suggest that carbonate weathering is the major player, with other factors playing a less significant role. The curve slightly lags conodont extinctions, hence the two events may not represent the same thing. Therefore, the term ''Lau event'' is used only for the extinction, not the following isotopic activity, which is named after the time period in which it occurred. A positive excursion of in pyrite coincides with the positive excursion following the Lau event, likely related to the expansion of euxinic conditions and enhanced pyrite burial.


Sedimentological impact

Profound sedimentary changes occurred at the beginning of the Lau event; these are probably associated with the onset of
sea level rise The sea level has been rising from the end of the last ice age, which was around 20,000 years ago. Between 1901 and 2018, the average sea level rose by , with an increase of per year since the 1970s. This was faster than the sea level had e ...
, which continued through the event, reaching a high point at the time of deposition of the Burgsvik beds, after the event. These changes appear to display
anachronism An anachronism (from the Greek , 'against' and , 'time') is a chronological inconsistency in some arrangement, especially a juxtaposition of people, events, objects, language terms and customs from different time periods. The most common type ...
, marked by an increase in erosional surfaces and the return of flat-pebbled conglomerates in the Eke beds. This is further evidence of a major blow to ecosystems of the time – such deposits can only form in conditions similar to those of the early
Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordov ...
period, when life as we know it was only just becoming established. Indeed,
stromatolite Stromatolites ( ) or stromatoliths () are layered Sedimentary rock, sedimentary formation of rocks, formations (microbialite) that are created mainly by Photosynthesis, photosynthetic microorganisms such as cyanobacteria, sulfate-reducing micr ...
s, which rarely form in the presence of abundant higher life forms, are observed during the Lau event and, occasionally, in the overlying Burgsvik beds; microbial colonies of '' Rothpletzella'' and '' Wetheredella'' become abundant. This suite of characteristics is common to the larger end-Ordovician and end-Permian extinctions.


See also

*
Anoxic event An anoxic event describes a period wherein large expanses of Earth's oceans were depleted of dissolved oxygen (O2), creating toxic, euxinic ( anoxic and sulfidic) waters. Although anoxic events have not happened for millions of years, the geol ...


Further reading


Linking the progressive expansion of reducing conditions to a stepwise mass extinction event in the late Silurian oceans


References

{{ExtEvent_nav Extinction events Silurian Isotope excursions