Larix Decidua

''Larix decidua'', the European larch, is a species of larch native to the mountains of central Europe, in the Alps and Carpathian Mountains, with small disjunct lowland populations in northern Poland. Its life span has been confirmed to be close to 1000 years, with ages of around 2000 years likely.

Description

''Larix decidua'' is a medium-size to large deciduous coniferous tree reaching 25–45 m tall, with a trunk up to 1.5 m diameter (exceptionally, to 53.8 m tall and 11.20 m girth .56 m diameterref name=MTCH>). The crown is conic when young, becoming broad and often irregular with age; the main branches are level to upswept, with the side branches often pendulous. The shoots are dimorphic, with growth divided into long shoots (typically 10–50 cm long) and bearing several buds, and short shoots only 1–2 mm long with only a single bud. The leaves are needle-like, light green, 2–4 cm long which turn bright yellow before they fall in the autumn, leaving the pale yellow-buff shoots bare until the next spring.

The cones are erect, ovoid-conic, 2–6 cm long, with 10-90 erect or slightly incurved (not reflexed) seed scales; they are bright pinkish-red at pollination, then green variably flushed red to purplish when immature, and turn brown and opening to release the seeds when mature, 4–6 months after pollination. The old cones commonly remain on the tree for many years, turning dull grey-black.

It is very cold tolerant, able to survive winter temperatures down to at least -50 °C, and is among the tree line trees in the Alps, reaching 2400 m altitude, though most abundant from 1000–2000 m. It only grows on well-drained soils, avoiding waterlogged ground and is not shade tolerant.

It is closely related to Siberian larch ''Larix sibirica'' and similar in morphology; the two are best distinguished by the dense pubescence on the cone scales of ''L. sibirica'', compared to the glabrous or only thinly pubescent cone scales of ''L. decidua''.

Subtaxa

The following varieties are accepted by ''Plants of the World Online'':
*''Larix decidua'' var. ''decidua'' – European larch or Alpine larch. Most of the range, except as below. Cones 2.5–6 cm; shoots yellow-buff.
*''Larix decidua'' var. ''carpatica'' Domin – Carpathian larch. In the Carpathian Mountains. Differs little from nominate var. ''decidua''.
*''Larix decidua'' var. ''polonica'' (Racib. ex Wóycicki) Ostenf. & Syrach – Polish larch. Disjunct in lowland northern and central Poland. Cones 2–3 cm; shoots very pale yellow-buff, almost white.
The Polish larch, being disjunct and growing in a different lowland habitat with a more continental rather than montane sub-oceanic climate, is widely treated at the higher rank of subspecies rather than variety, ''Larix decidua'' subsp. ''polonica'' (Racib. ex Wóycicki) Domin. The Russian botanist Evgenij Bobrov considered it to be an ancient natural hybrid between European larch and Siberian larch ''L. sibirica'', but this has been shown to be incorrect, with it derived from only ''L. decidua''.

Ecology

The seeds are an important food for some birds, notably siskin, redpoll, and citril finch, while the buds and immature cones are eaten by capercaillie.

Larch needles (including ''L. decidua'' and ''L. sibirica'') are the only known food for caterpillars of the case-bearer moth '' Coleophora sibiricella''; its cone scales are used as food by the caterpillars of the tortrix moth '' Cydia illutana''.

It is susceptible to larch canker '' Lachnellula willkommii''.

Cultivation

''Larix decidua'' is widely cultivated as an ornamental tree for planting in gardens and parks. It was first cultivated in Britain in or shortly before 1629. John Evelyn encouraged its wider planting and use. Three successive Dukes of Atholl planted it widely and the fourth Duke wrote "Observations on Larch" in 1807 further encouraged its cultivation, which he practiced on a large scale. One of the larches planted by the second Duke of Atholl at Dunkeld in 1737 is still standing, as are two slightly older specimens planted in 1725 at Kailzie, near Peebles in southern Scotland.

European larch is now widely naturalised in northern Europe, including Britain, Scandinavia, and in Germany north of its native range. It is also widely cultivated in southern Canada and the northeastern United States, and is naturalised in Maine, Michigan, New York, Pennsylvania, Connecticut, New Hampshire, Vermont, and Rhode Island. In the northern Appalachian Mountains it is often used for the reforestation of surface mines. European larch can grow on drier soils and tolerate warmer climates than tamarack (''Larix laricina'') or Siberian larch (''Larix sibirica''), being better suited to non-boreal climates.

Hybrids

European larch readily hybridises with Japanese larch ''L. kaempferi'' when the two are planted together; the hybrid, named Dunkeld larch or hybrid larch ('' Larix × marschlinsii'' Coaz, syn. ''L. × eurolepis''), is extensively used in forestry, and is also naturalised in Britain, Ireland, and elsewhere. It is faster-growing than either parent, and more resistant to larch canker '' Lachnellula willkommii''. Hybrids with several other larches have been made, but are rare; the hybrid with ''Larix laricina'' has also been named, as ''Larix × pendula'' (Solander) Salisbury.

Bonsai

The European larch is a popular bonsai species, with many unique specimens available in European circles, and is popularly used in bonsai forest groups.

Invasive species

In New Zealand, ''Larix decidua'' is classed as a wilding conifer, an invasive species which spreads into the high country, where it had been planted by the New Zealand Forest Service for erosion control.

Uses

The wood is tough and durable, but also flexible in thin strips, and is particularly valued for yacht building; wood used for this, known as 'boatskin larch', must be free of knots, and can only be obtained from old trees that were pruned when young to remove side branches. Small larch poles are widely used for rustic fencing.

Because of its fast juvenile growth and its pioneer character, larch has found numerous applications in forestry and agroforestry. It is used as a 'preparatory species' to afforest open land, abandoned farmland or disturbed land, and as a 'nurse species' prior to the introduction of more demanding species.

Gallery

File:Larix decidua Aletschwald.jpg, ''Larix decidua'' subsp. ''decidua'' at Aletschwald, Switzerland
File:Larix decidua ssp polonica, Gajówka, Kraków, Poland.jpg, ''Larix decidua'' subsp. ''polonica'' at Gajówka, Kraków, Poland
File:Larch in Guarda (crop).jpg, In winter, without leaves, Guarda, Graubünden, Switzerland
File:Albeck Seebachern bizarre Laerche 25102013 872.jpg, Larch in Albeck, district Feldkirchen, Carinthia, Austria
File:ArmentaraFels.jpg, South Tyrol, Wengen-La Val-La Valle
File:Larix decidua Embrun1.jpg, Parc régional du Queyras, Embrun, Hautes-Alpes, France
File:Albeck Seebachern Laerchenwald 25102013 869.jpg, Autumnal forest with larches, spruces and arolla pines at Seebachern
File:Bergtocht van Peio Paese naar Lago Covel in het Nationaal park Stelvio (Italië). Wond van een afgebroken tak. Europese larix (Larix decidua) 02.jpg, Bark, with a wound from a broken branch.
File:Larix decidua0.jpg, European larch foliage and cones.
File:Larix decidua developing cone.JPG, Developing seed cone in detail.
File:Larix-decidua 0088 S.jpg, Young seed cones (red) and pollen cones (yellow).
File:LarixDeciduaFemaleFlower.jpg, Young female cone
File:Larix decidua male cones 2.jpg, Male cones
File:Larch planted on a scree slope.JPG, European larch planted for erosion control on a scree slope in Canterbury, New Zealand.

References

External links

''Larix decidua''
Information, genetic conservation units and related resources. European Forest Genetic Resources Programme (EUFORGEN)

{{Authority control

decidua
Trees of Europe
Flora of France
Flora of the Alps
Flora of Austria
Flora of the Czech Republic
Flora of Germany
Flora of Italy
Flora of Poland
Flora of Romania
Flora of Switzerland
Flora of Ukraine
Least concern plants
Trees of humid continental climate
Garden plants of Europe
Ornamental trees
Deciduous conifers
Taxa named by Philip Miller
Flora of the Carpathians