__NOTOC__ ''Laornis'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of a prehistoric neornithine

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s, known only from Specimen YPM 820, a single

tibiotarsus The tibiotarsus is the large bone between the femur and the tarsometatarsus in the leg of a bird. It is the fusion of the proximal part of the tarsus with the tibia. A similar structure also occurred in the Mesozoic Heterodontosauridae. These ...

leg bone discovered in the late 19th century. Consequently, the genus is

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

, containing only the

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

''Laornis edvardsianus''. Regarding its

scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

, ''Laornis'' means "stone bird", from

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

''lao'' "stone" + ''ornis'' "bird". ''edvardsianus'' honors

Alphonse Milne-Edwards Alphonse Milne-Edwards (Paris, 13 October 1835 – Paris, 21 April 1900) was a French mammalogist, ornithologist, and carcinologist. He was English in origin, the son of Henri Milne-Edwards and grandson of Bryan Edwards, a Jamaican planter who ...

, to compliment the French

paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...

on his landmark study ''Recherches Anatomiques et Paleontologiques pour servir a l'Histoire des Oiseaux Fossiles de la France'', the second part of which was nearing completion at that time. It was found in Late

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...

or Early

Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...

sediments of the

Hornerstown Formation The Hornerstown Formation is a latest Cretaceous to early Paleocene-aged geologic Formation (geology), formation in New Jersey. It preserves a variety of fossil remains, including those of dinosaurs, and contains direct evidence of the Mass morta ...

at the Birmingham

Marl Marl is an earthy material rich in carbonate minerals, Clay minerals, clays, and silt. When Lithification, hardened into rock, this becomes marlstone. It is formed in marine or freshwater environments, often through the activities of algae. M ...

Pits,

Pemberton Township, New Jersey Pemberton Township is a Township (New Jersey), township in Burlington County, New Jersey, Burlington County, in the U.S. state of New Jersey. As of the 2020 United States census, the township's population was 26,903, a decrease of 1,009 (−3.6% ...

, United States (39°59'N, 74°43'W). The deposits were laid down at about 66–63 Ma (

million years ago Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used w ...

). The bone is rather distinct but not very diagnostic. Its general shape suggests that ''Laornis'' was a semi-aquatic bird with longish legs and a body at least the size of a large goose. It may have been a wading bird, in which case it stood probably around one meter (3–4 ft) tall in life, depending on how long its legs and neck were exactly, which of course cannot be told from the one known bone. On the other hand, it might have been a larger

seabird Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...

with proportionally shorter legs. It has variously been allied with the

Charadriiformes Charadriiformes (, from '' Charadrius'', the type genus of family Charadriidae) is a diverse order of small to medium-large birds. It includes about 390 species and has members in all parts of the world. Most charadriiform birds live near water ...

and the

Gruiformes The Gruiformes ( ) are an order containing a considerable number of living and extinct bird families, with a widespread geographical diversity. Gruiform means "crane-like". Traditionally, a number of wading and terrestrial bird families that ...

, and is tentatively placed in a family of its own, Laornithidae. It may be considered some kind of basal gruiform, or more probably part of an ancestral lineage related to the common ancestor of gruiform, charadriiform, and/or any or all other modern "wading" bird families. It might have been one of the extinct stilt-legged

waterfowl Anseriformes is an order of birds also known as waterfowl that comprises about 180 living species of birds in three families: Anhimidae (three species of screamers), Anseranatidae (the magpie goose), and Anatidae, the largest family, which i ...

of the

Presbyornithidae Presbyornithidae is an extinct group of birds found in North America, South America, East Asia, Australia and possibly North Africa. They had evolved by the Maastrichtian age of the Late Cretaceous and became extinct during the Aquitanian age of ...

, and it cannot even be excluded that it was an ancient

pseudotooth bird The Pelagornithidae, commonly called pelagornithids, pseudodontorns, bony-toothed birds, false-toothed birds or pseudotooth birds, are a prehistoric family of large seabirds. Their fossil remains have been found all over the world in rocks dating ...

, seabirds of unclear affiliation that

evolved Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...

to immense proportions in the

Neogene The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of th ...

but by the time of ''Laornis'' probably were mostly the size of a large

petrel Petrels are tube-nosed seabirds in the phylogenetic order Procellariiformes. Description Petrels are a monophyletic group of marine seabirds, sharing a characteristic of a nostril arrangement that results in the name "tubenoses". Petrels enco ...

References

External links

Topographic map of the type locality
from

TopoQuest TopoQuest is a free web mapping service built on open source software that provides internet-based topographic map for most of the United States. The site is one of three internet services used by Wikipedia for providing topographic maps. It arose ...

{{Taxonbar, from=Q6487938 Birds Bird genera Paleogene birds Taxa named by Othniel Charles Marsh Fossil taxa described in 1870 Birds of North America