Lanthanosuchoidea is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

superfamily SUPERFAMILY is a database and search platform of structural and functional annotation for all proteins and genomes. It classifies amino acid sequences into known structural domains, especially into SCOP superfamilies. Domains are functional, str ...

of ankyramorph

parareptile Parareptilia ("near-reptiles") is an extinct group of Basal (phylogenetics), basal Sauropsida, sauropsids ("Reptile, reptiles"), traditionally considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds ...

s from the middle

Pennsylvanian Pennsylvanian may refer to: * A person or thing from Pennsylvania * Pennsylvanian (geology) The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, on the International Commission on Stratigraphy, ICS geologic timesc ...

to the middle

Guadalupian The Guadalupian is the second and middle Series (stratigraphy), series/Epoch (geology), epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico an ...

epoch In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured. The moment of epoch is usually decided b ...

(

Moscovian Moscovian may refer to: *An inhabitant of Moscow, the capital of Russia *Something of, from, or related to Moscow *Moscovian (Carboniferous) The Moscovian is in the International Commission on Stratigraphy, ICS geologic timescale a stage (strat ...

Wordian In the geologic timescale, the Wordian is an age or stage of the Permian. It is the middle of three subdivisions of the Guadalupian Epoch or Series. The Wordian lasted between and million years ago (Ma). It was preceded by the Roadian and fo ...

stages) of

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

and

Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...

. It was named by the Russian paleontologist Ivachnenko in 1980, and it contains two families

Acleistorhinidae Acleistorhinidae is an extinct family of Late Carboniferous and Early Permian-aged ( Moscovian to Kungurian stage) parareptiles. It is defined as a node based clade including the last common ancestor of '' Acleistorhinus pteroticus'' and '' Co ...

and

Lanthanosuchidae Lanthanosuchidae is a family of procolophonomorph parareptile Parareptilia ("near-reptiles") is an extinct group of Basal (phylogenetics), basal Sauropsida, sauropsids ("Reptile, reptiles"), traditionally considered the sister taxon to Eurept ...

Phylogeny

Lanthanosuchoidea is a

node-based taxon Phylogenetic nomenclature is a method of nomenclature for taxon, taxa in biology that uses phylogenetics, phylogenetic definitions for taxon names as explained below. This contrasts with Biological classification, the traditional method, by which ...

defined in 1997 as "the most recent common ancestor of ''

Lanthanosuchus ''Lanthanosuchus'' is an extinct genus of parareptile from the Late Permian. It was found at Isheevo in Tatarstan Tatarstan, officially the Republic of Tatarstan, sometimes also called Tataria, is a Republics of Russia, republic of Russia ...

'', ''

Lanthaniscus ''Lanthaniscus'' is an extinct genus of lanthanosuchoid ankyramorph parareptile known from the Guadalupian epoch (Late Roadian to latest Wordian age) of Eastern Europe, Russia. ''Lanthaniscus'' was first named by M. F. Ivakhnenko in 1980 and th ...

'', and ''

Acleistorhinus ''Acleistorhinus'' (ah-kles-toe-RYE-nuss) is an extinct genus of parareptile known from the Early Permian (middle Kungurian stage) of Oklahoma. It is notable for being the earliest known anapsid reptile yet discovered. The morphology of the lower ...

''". The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below follows the topology from a 2011 analysis by Ruta ''et al.'' The

below follows the topology from a 2016 analysis by MacDougall ''et al.'' However, the phylogenetic analysis conducted by Cisneros ''et al.'' (2021) did not recover a clade uniting lanthanosuchids and acleistorhinids to the exclusion of all other parareptiles. Instead, acleistorhinids were recovered as the

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

of the clade

Procolophonia Procolophonia is an extinct suborder (clade) of herbivorous reptiles that lived from the Middle Permian till the end of the Triassic period. They were originally included as a suborder of the Cotylosauria (later renamed Captorhinida Carroll ...

, while lanthanosuchids were recovered within the procolophonian subgroup

Pareiasauromorpha Pareiasauromorpha is a group of parareptilian amniotes from the Permian. It includes genera found all over the world, with many genera from Asia and South Africa. The clade was first used as a group by Linda A. Tsuji in 2011, in order to group t ...

References

Procolophonomorpha Permian reptiles Prehistoric reptiles of Asia Prehistoric reptiles of Europe Prehistoric reptiles of North America {{Permian-reptile-stub