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''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s belonging to
Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark ...
containing over 100
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
. In the present circumscription, it contains 6 tribes (
Tritomini Tritomini is a tribe of pleasing fungus beetles in the family Erotylidae. There are about 7 genera and at least 30 described species in Tritomini. Genera These genera belong to the tribe Tritomini: * '' Callischyrus'' * ''Haematochiton ' ...
, Dacnini, Megalodacnini, Erotylini, Cryptophilini, and
Languriini Languriini is a tribe of beetles, known as lizard beetles, in the family Erotylidae, subfamily Languriinae. Lizard beetles are commonly found on leaves or flowers, and their larvae bore into plant stems. Adults are generally long and parallel-sid ...
) and 10 subfamilies ( Cryptophilinae, Dacninae, Encaustinae,
Erotylinae Erotylinae is a subfamily of pleasing fungus beetle, pleasing fungus beetles in the family Erotylidae. Selected genera * ''Acropteroxys'' Gorham, 1887 * ''Antillengis'' * ''Aegithus'' * ''Altisessor'' * ''Aulacochilus'' * ''Bacis (beetle), Baci ...
,
Languriinae Languriinae is a subfamily of pleasing fungus beetle in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present ...
, Loberinae, Megalodacninae, Pharaxonothinae, Tritominae, and
Xenoscelinae Xenoscelinae is a subfamily of pleasing fungus beetles in the family Erotylidae. There are about 8 genera and 12 described species in Xenoscelinae. Genera These eight genera belong to the subfamily Xenoscelinae: * '' Cryptophilus'' Reitter, 187 ...
). In other words, the narrowly circumscribed Erotylidae correspond to the subfamily Erotylinae in the definition ''
sensu lato ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular co ...
''. There are doubts on the
monophyly In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent comm ...
of lower ranked taxa within Erotylidae, with further phylogenetic studies requiring better sampling and studies of unexplored character sets, for example the metendosternite and penile flagellum, which are generally lacking detailed morphological studies within the Coleoptera literature. The Eroytlina taxonomy is based on traits such as their different colors and not off morphological differences like mouthparts, thorax, and abdominal terminalia (Pecci-Maddalena). Erotylidae feed on
plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
and
fungal A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as one of the tradit ...
matter; some are important
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female carpel, stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are ...
s (e.g. of the ancient
cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...
s), while a few have gained notoriety as pests of some significance. Sometimes, useful and harmful species are found in one
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
, e.g. '' Pharaxonotha''. Most pleasing fungus beetles, however, are inoffensive animals of little significance to humans. The oldest fossil is an undescribed species known from Early Cretaceous (
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
)
Lebanese amber Lebanese amber is fossilized resin found in Lebanon and its surroundings. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropic ...
.


Selected genera

These 160 genera belong to the family Erotylidae: * '' Acropteroxys'' Gorham, 1887 i c g b * '' Acryptophagus'' * '' Aegithus'' Fabricius, 1801 g * '' Amblyopus'' Lacordaire, 1842 g * '' Amblyscelis'' Gorham, 1888 * '' Anadastus'' Gorham, 1887 g * '' Apolybas'' Alvarenga, 1965 g * '' Atomarops'' Reitter * '' Aulacochilus'' Lacordaire, 1842 * ''
Bacis Bakis (also Bacis; ) is a general name for the inspired prophets and dispensers of oracles who flourished in Greece from the 8th to the 6th century B.C. Philetas of Ephesus,Suda s. v. Βάκις Aelian and John Tzetzes distinguish between three: ...
'' Dejean, 1836 g * '' Barytopus'' Chevrolat, 1836 g * '' Bolerus'' Grouvelle * '' Brachypterosa'' Zablotny & Leschen, 1996 * '' Brachysphaenus'' * '' Caenolanguria'' Gorham, 1887 g * '' Callischyrus'' * '' Camptocarpus'' * '' Cathartocryptus'' Sharp, 1886 i c g * '' Chinophagus'' Ljubarsky, 1997 * '' Cladoxena'' Motschulsky * '' Cnecosa'' * '' Coccimorphus'' Hope, 1841 * '' Coelocryptus'' Sharp, 1900 * '' Combocerus'' Bedel, 1868 g * '' Coptengis'' Crotch, 1876 * '' Crotchia'' Fowler * '' Crowsenguptus'' * '' Cryptodacne'' Sharp, 1878 g * '' Cryptophilus'' Reitter, 1874 i c g b * '' Cycadophila'' Xu, Tang & Skelley, 2015 g * '' Cypherotylus'' Crotch, 1873 * '' Cyrtomorphus'' Lacordaire, 1842 g * '' Cytorea'' * '' Dacne'' Latrielle, 1796 * '' Dactylotritoma'' Arrow, 1925 g * '' Dapsa'' * '' Dasydactylus'' Gorham, 1887 i c g b * '' Doubledaya'' White, 1850 g * '' Ectrapezidera'' * '' Ellipticus'' Chevrolat, 1836 g * '' Empocryptus'' Sharp * '' Encaustes'' Lacordaire, 1842 g * '' Epilanguria'' Fowler, 1908 g * '' Episcapha'' Dejean, 1837 * '' Episcaphula'' Crotch * '' Erotylina'' Curran, 1944 g * '' Erotylus'' Fabricius, 1775 * '' Eutriplax'' Lewis, 1887 * '' Fitoa'' * '' Goniolanguria'' * ''
Haematochiton ''Haematochiton'' is a genus of pleasing fungus beetles in the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the pres ...
'' Gorham, 1888 i c g b * '' Hapalips'' Reitter, 1877 i c g b * '' Henoticonus'' Reitter, 1878 * '' Hirsotriplax'' * '' Hirsutotriplax'' Skelley, 1993 i c g b * '' Homoeotelus'' Hope, 1841 * '' Hoplepiscapha'' Lea, 1922 * '' Hornerotylus'' g * ''
Iphiclus ''Iphiclus'' is a genus of beetles in the family Erotylidae. It was first described by Louis Alexandre Auguste Chevrolat Louis Alexandre Auguste Chevrolat (29 March 1799, Paris – 16 December 1884, Paris) was a French entomologist. He speci ...
'' Chevrolat, 1836 g * '' Ischyrus'' Lacordaire, 1842 i c g b * '' Languria'' Latreille, 1802 i c g b * '' Languriomorpha'' * '' Langurites'' Motschulsky, 1860 i c g b * '' Lepidotoramus'' Leschen, 1997 i c g * '' Leucohimatium'' Rosenhauer, 1856 * '' Ligurana'' Chûjô, 1974 g * '' Linodesmus'' Bedel, 1882 * '' Loberogosmus'' Reitter * '' Loberolus'' * '' Loberonotha'' Sen Gupta and Crowson, 1969 i c g * '' Loberopsyllus'' * '' Loberoschema'' Reitter * '' Loberus'' LeConte, 1861 i c g b * '' Lobosternum'' Reitter, 1875 i c g * '' Lybanodes'' * '' Lybas'' Lacordaire, 1842 g * '' Macromelea'' Hope * '' Macrophagus'' Motschulsky * '' Malleolanguria'' * '' Megalodacne'' Crotch, 1873 i c g b * '' Megischyrus'' Crotch, 1873 g * '' Meristobelus'' * '' Micrencaustes'' Crotch, 1875 g * '' Microlanguria'' Lewis * ''
Microsternus ''Microsternus'' is a genus of pleasing fungus beetles in the family Erotylidae. Species These three species belong to the genus ''Microsternus'': * '' Microsternus perforatus'' (Lewis, 1883) * '' Microsternus tricolor'' Lewis, 1887 * '' Micro ...
'' Lewis, 1887 i c g b * '' Mimodacne'' * '' Mycetaea'' Stephens, 1829 * '' Mycolybas'' Crotch, 1876 * '' Mycophtorus'' Lacordaire, 1842 g * '' Mycotretus'' Lacordaire, 1842 i c g b * '' Neocladoxena'' Maeda, 1974 g * '' Neodacne'' Chûjô, 1976 g * '' Neoloberolus'' Leschen, 2003 i c g * '' Neopriotelus'' Alvarenga, 1965 g * '' Neosternus'' Dai & Zhao, 2013 g * '' Neotriplax'' Lewis, 1887 * '' Neoxestus'' Crotch, 1875 g * '' Nomotus'' * '' Oligocorynus'' Chevrolat, 1836 g * '' Oretylus'' Heller, 1920 g * '' Ortholanguria'' * '' Othniocryptus'' Sharp, 1900 * '' Pachylanguria'' Crotch, 1875 g * '' Paederolanguria'' Mader, 1939 g * '' Paphezia'' Zablotny & Leschen, 1996 * '' Paracladoxena'' Fowler * '' Paraxonotha'' g * ''
Pediacus ''Pediacus'' is the largest genus in the family Cucujidae of flat bark beetles. It contains 31 currently recognized species.Michael C. Thomas. 2004. A revision of ''Pediacus'' Shuckard (Coleoptera: Cucujidae) for America north of Mexico, with no ...
'' * '' Penolanguria'' Kolbe * '' Pharaxonotha'' Reitter, 1875 i c g b * '' Phricobacis'' Crotch, 1876 g * '' Platoberus'' Sharp * '' Prepopharus'' Erichson, 1847 g * '' Promecolanguria'' * '' Protoloberus'' Leschen, 2003 i c g * '' Pselaphacus'' Percheron, 1835 g * '' Pselaphandra'' Jacobson, 1904 * '' Pseudhapalips'' Champion * '' Pseudhenoticus'' Sharp * '' Pseudischyrus'' Casey, 1916 i c g b * '' Pseudotritoma'' Gorham, 1888 g * '' Rhodotritoma'' Arrow, 1925 g * '' Scaphengis'' Gorham, 1888 g * ''
Scaphidomorphus ''Scaphidomorphus'' is a genus of beetles belonging to the family Erotylidae ''Ischyrus quadripunctatus'' Erotylidae, or the pleasing fungus beetles, is a family of beetles belonging to Cucujoidea containing over 100 genera. In the present ...
'' Hope, 1841 g * '' Scaphodacne'' Heller, 1918 * '' Scelidopetalon'' Delkeskamp, 1957 g * '' Setariola'' Jakobson, 1915 g * '' Sphenoxus'' Lacordaire, 1842 g * '' Spondotriplax'' Crotch, 1875 g * '' Stengita'' * '' Stenodina'' * '' Strongylosomus'' Chevrolat, 1836 g * '' Tapinotarsus'' Kirsch, 1865 g * '' Telmatoscius''* '' Teretilanguria'' * '' Tetralanguria'' Crotch, 1875 g * '' Tetraphala'' g * '' Tetratritoma'' Arrow, 1925 g * '' Thallis'' * '' Thallisella'' Crotch * '' Thallisellodes'' Arrow, 1925 g * '' Tomarops'' Grouvelle, 1903 * '' Toramus'' Grouvelle, 1916 i c g b * '' Trapezidera'' * '' Trapezidistes'' Fowler, 1887 g * '' Trichotritoma'' * '' Trichulus'' * '' Triplacidea'' Gorham, 1901 * ''
Triplax ''Triplax'' is a genus of pleasing fungus beetles in the family Erotylidae. There are about 18 described species in ''Triplax''. Species * '' Triplax alachuae'' Boyle, 1956 * '' Triplax californica'' LeConte, 1854 * '' Triplax cuneata'' Boyle, 19 ...
'' Herbst, 1793 i c g b * '' Tritoma'' Fabricius, 1775 i c g b * '' Truquiella'' b * '' Typocephalus'' Chevrolat, 1837 g * '' Xenocryptus'' Arrow, 1929 * '' Xenohimatium'' Lyubarsky & Perkovsky, 2012 g * '' Xenoscelinus'' Grouvelle, 1910 g * '' Xenoscelis'' Wollaston, 1864 g * '' Xestus'' Wollaston, 1864 g * '' Zavaljus'' Reitter, 1880 g* '' Acropteroxys'' Gorham, 1887 * '' Zonarius'' (= ''Oligocorynus'') * '' Zythonia'' Westwood, 1874 Data sources: i = ITIS, c = Catalogue of Life, g = GBIF, b = Bugguide.net


References

* * * *


External links

* *
Key to the British species of family Erotylidae
{{Authority control Cucujoidea families