Lacertoidea
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The Lacertoidea is a group of
squamate Squamata (, Latin ''squamatus'', 'scaly, having scales') is the largest Order (biology), order of reptiles; most members of which are commonly known as Lizard, lizards, with the group also including Snake, snakes. With over 11,991 species, it i ...
reptiles that includes the
Lacertidae The Lacertidae are the family of the wall lizards, true lizards, or sometimes simply lacertas, which are native to Afro-Eurasia. It is a diverse family with at about 360 species in 39 genera. They represent the dominant group of reptiles found ...
,
Teiidae Teiidae is a family of lacertoidean lizards native to the Americas. Members of this family are generally known as whiptails or racerunners; however, tegus also belong to this family. Teiidae is sister to the Gymnophthalmidae, Gymnopthalmidae, and ...
,
Gymnophthalmidae Gymnophthalmidae is a family of lizards with at least 250 species, sometimes known commonly as spectacled lizards or microteiids. They are called "spectacled" because of their transparent lower eyelids, which allow them to still see with closed ...
, and
Amphisbaenia Amphisbaenia (called amphisbaenians or worm lizards) is a group of typically legless lizards, comprising over 200 extant species. Amphisbaenians are characterized by their long bodies, the reduction or loss of the limbs, and rudimentary eyes. A ...
. The finding from molecular phylogenetic studies that the burrowing Amphisbaenia were nested in a clade with the
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
forms led Vidal & Hedges (2005) to propose a new name for the group based on shared morphogical characters, Laterata, "referring to the presence of tile-like (squarish or quadrangular, and sometimes raised) scales that form the rings in Amphisbaenia, and are also present ventrally in Lacertiformata and Teiformata". Studies of anatomy have traditionally grouped the lacertoids with
skink Skinks are a type of lizard belonging to the family (biology), family Scincidae, a family in the Taxonomic rank, infraorder Scincomorpha. With more than 1,500 described species across 100 different taxonomic genera, the family Scincidae is one o ...
s; however, more recent studies focusing on DNA have placed them as a distinct group of lizards, more closely related to the venomous
Toxicofera Toxicofera (Latin for "toxin-bearers") is a proposed clade of squamata, scaled reptiles (squamates) that includes the snake, Serpentes (snakes), Anguimorpha (monitor lizards, Helodermatidae, beaded lizards, and alligator lizards) and Iguania (igu ...
.Wiens, J. J., Hutter, C. R., Mulcahy, D. G., Noonan, B. P., Townsend, T. M., Sites, J. W., & Reeder, T. W. (2012). Resolving the phylogeny of lizards and snakes (Squamata) with extensive sampling of genes and species. Biology Letters, 8(6), 1043-1046. The relationships of the Amphisbaenia to other lizards have long been a mystery; although superficially snakelike, their anatomy and locomotion is quite distinct from that of snakes, Amphisbaenia have a reduced right lung as opposed to snakes which have their left lung reduced.Gans, C. (1978) The characteristics and affinities of the Amphisbaenia. Trans. Zool. Soc. Lond. (1978) 34:347-416. However, they do not closely resemble any other group of lizards. Recent molecular studies suggest that amphisbaenians are grouped with the Lacertidae, in a group named the Lacertibaenia. Lacertoids have a wide geographic distribution. The Lacertidae are found throughout Europe and Asia, with a major radiation endemic to Africa. The Teiidae are diverse in South America, but some members of the group occur in southern North America as well. The Amphisbaenia are especially widespread, occurring in North America, Europe, Africa, South America, and the Caribbean. The origins of the group are unclear. The oldest definitive lacertoid is the amphisbaenian '' Plesiorhineura'', from the early Paleocene of North America. Given that it resembles modern worm lizards, the lacertoids most likely appeared and diversified in the Cretaceous. However,
Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...
taxon ''
Purbicella ''Purbicella'' is a genus of extinct squamate from the Early Cretaceous of southern England ( Lulworth Formation). The type and only species is ''Purbicella ragei'', which was described by Susan E. Evans, Marc E. H. Jones, and Ryoko Matsumoto in ...
'' may be a lacertoid, and would be the oldest known. Within the Laterata there are many species listed on IUCN Redlist conservation status. This family listed 5 species extinct, 31 species critically endangered, 53 species endangered, 40 species vulnerable, 53 species near threatened, and 569 species as least concerned.


References

{{Taxonbar, from=Q13518423 Vertebrate superfamilies Selandian first appearances Taxa named by Nicolaus Michael Oppel Extant Selandian first appearances